%PDF-1.2 % 2 0 obj << /Length 9953 >> stream BT /F4 1 Tf 9 0 0 9 65.09 43.042 Tm 0 0 0 1 k /GS2 gs -0.0058 Tc 0.0613 Tw [(*Corresponding )47.9(Author E-mail: st3@soton.ac.uk)]TJ 0 -2.1323 TD -0.0053 Tc 0.0343 Tw [(\251 2001 )48.1(Association of )51.2(Applied Biologists)]TJ /F5 1 Tf 8 0 0 8 65.147 797.596 Tm -0.0033 Tc 0.0771 Tw (Ann. appl. Biol. )Tj /F6 1 Tf 6.7323 0 TD -0.0056 Tc (\(2001\))Tj /F5 1 Tf 2.6362 0 TD -0.0031 Tc (, )Tj /F7 1 Tf 0.5811 0 TD -0.0087 Tc (139)Tj /F6 1 Tf 1.4811 0 TD -0.008 Tc [(:1)27.4(1)38(1-1)27.4(18)]TJ /F5 1 Tf -11.4307 -1.1232 TD -0.0045 Tc 0.0665 Tw [(Printed in Gr)34.5(eat Britain)]TJ /F4 1 Tf 9 0 0 9 522.205 789.035 Tm -0.0019 Tc (111)Tj /F8 1 Tf 14 0 0 14 92.444 761.001 Tm -0.005 Tc 0.0294 Tw [(Resour)13.2(ce pr)11.2(ovision for)13.2( farmland gamebirds: the value of beetle banks)]TJ /F9 1 Tf 11 0 0 11 174.536 735.999 Tm -0.0071 Tc 0.0253 Tw [(By S R )11(THOMAS*, D GOULSON and J M HOLLAND)]TJ /F10 1 Tf 6.4 0 0 6.4 423.106 739.656 Tm 0 Tc 0 Tw (1)Tj /F11 1 Tf 11 0 0 11 75.89 711.99 Tm -0.0056 Tc 0.0607 Tw (Biodiversity and Ecology Division, School of Biological Sciences, University of Southampton, Bassett)Tj 9.1276 -1.09 TD -0.0059 Tc 0.0484 Tw [(C)9.7(r)30.2(escent East, Southampton, Hampshir)30.2(e SO16 7PX, UK)]TJ /F10 1 Tf 6.4 0 0 6.4 140.548 691.637 Tm 0 Tc 0 Tw (1)Tj /F11 1 Tf 11 0 0 11 143.752 688.009 Tm -0.0065 Tc 0.0481 Tw [(The Game Conser)-6.5(vancy T)50.2(r)1.3(ust, For)27(dingbridge, Hampshir)29.6(e SP6 1EF)119.8(, UK)]TJ /F9 1 Tf 3.5588 -2.1827 TD 0 Tc 0 Tw (\()Tj /F11 1 Tf 0.3273 0 TD -0.0058 Tc 0.0447 Tw [(Accepted 20 )12.1(April 2001; Received 17 January 2001\))]TJ /F12 1 Tf 8.2179 -2.3141 TD 0.0008 Tc (Summary)Tj /F9 1 Tf -17.1625 -2.1827 TD -0.0106 Tc -0.0613 Tw [(Severe declines have occurred in the populations of wild game birds in Britain. )12.7(This has been attributed)]TJ -0.8246 -1.09 TD -0.0056 Tc -0.0328 Tw (to agricultural intensification, leading to the loss of invertebrates vital within chick diets, fewer feeding)Tj T* -0.0472 Tw [(resources for adults, and inadequate provision of nesting and brood-rearing habitat. )12.5(This paper explores)]TJ 0 -1.0926 TD -0.004 Tc 0.1227 Tw (the potential value of simple sown grass strips \226 beetle banks \226 in providing these resources, and)Tj 0 -1.09 TD -0.0061 Tc 0.1045 Tw [(compares results with functionally similar conventional field mar)11.9(gins. )11.9(The data indicate that beetle)]TJ T* -0.0058 Tc 0.1009 Tw [(banks can contribute useful, albeit lower)32.9(, densities of chick-food than conventional mar)12.2(gins. )15(These)]TJ 0 -1.0926 TD -0.0285 Tw (resources are more abundant later in the season, which may have implications for early hatched chicks.)Tj 0 -1.09 TD -0.0055 Tc -0.039 Tw (Beetle banks provide considerable quantities of nesting cover for adults, although sheltering conditions)Tj T* -0.011 Tc -0.0613 Tw (may never be as satisfactory as in well managed hedgerows. Given the ease and low cost of establishment)Tj T* -0.0061 Tc -0.038 Tw (of beetle banks, we suggest that they may be valuable components within a range of game management)Tj 0 -1.0926 TD -0.0057 Tc 0.0363 Tw [(techniques on the farm, as a \221spin-of)9.8(f)-54.6(\222)71.7( to their primary role as overwintering habitat for polyphagous)]TJ 0 -1.09 TD -0.006 Tc 0.0479 Tw [(predators. )9.5(They may be important particularly where resources for game birds are impoverished, but)]TJ T* 0.0539 Tw [(clearly cannot substitute for suitably managed field mar)9.5(gins.)]TJ /F12 1 Tf 0 -3.2727 TD -0.0255 Tc 0.1228 Tw (Key words:)Tj /F9 1 Tf 4.7931 0 TD -0.0235 Tc 0.1225 Tw [( Field mar)7.4(gin, chick-food invertebrates, nesting cover)35.8(, biodiversity)59(, agroecosystems, arable)]TJ /F12 1 Tf 1.4044 -2.6388 TD -0.0001 Tc [(Intr)10.2(oduction)]TJ /F11 1 Tf 0.3891 -2.1801 TD -0.007 Tc 0.0637 Tw [(Game bir)29.1(d)6(s)]TJ /F9 1 Tf -7.0273 -1.09 TD -0.006 Tc 0.1137 Tw (Grey partridge \()Tj /F11 1 Tf 6.579 0 TD -0.0081 Tc 0.1152 Tw [(Per)28(dix per)30.6(dix)]TJ /F9 1 Tf 5.4296 0 TD -0.0057 Tc 0.1159 Tw (\) populations have)Tj -12.8306 -1.0926 TD 0.0013 Tc 0.1228 Tw (declined drastically over the last few decades. In)Tj 0 -1.09 TD -0.0055 Tc 0.042 Tw (the 1950s, average densities of around 25 pairs per)Tj T* -0.0182 Tc (km)Tj /F10 1 Tf 6.4 0 0 6.4 79.008 337.193 Tm -0.0043 Tc (-2)Tj /F9 1 Tf 11 0 0 11 84.309 333.565 Tm -0.0057 Tc 0.0444 Tw [( could be found, whereas fewer than five pairs)]TJ -1.742 -1.0926 TD 0.0001 Tc 0.1227 Tw (per km)Tj /F10 1 Tf 6.4 0 0 6.4 97.462 325.202 Tm 0.0046 Tc (-2)Tj /F9 1 Tf 11 0 0 11 102.904 321.546 Tm 0.0033 Tc [( were recorded by the mid 1980s \(Potts,)]TJ -3.4325 -1.0901 TD -0.0059 Tc 0.0802 Tw [(1986\). Consequently)55.9(, this species was entered into)]TJ T* -0.0062 Tc 0.0576 Tw (the UK Red Data Book \(Batten )Tj /F11 1 Tf 12.9466 0 TD -0.0063 Tc 0.0545 Tw (et al.)Tj /F9 1 Tf 2.0229 0 TD -0.0054 Tc 0.0591 Tw (, 1990\). Only)Tj -14.9695 -1.0926 TD -0.0061 Tc 0.107 Tw [(lar)11.9(ge-scale reared bird releases are responsible for)]TJ 0 -1.09 TD 0.0461 Tc 0.1227 Tw [(maintaining stable populations of red-legged)]TJ T* -0.0059 Tc -0.0318 Tw (partridge \()Tj /F11 1 Tf 4.0999 0 TD -0.0064 Tc -0.0363 Tw (Alectoris rufa)Tj /F9 1 Tf 5.409 0 TD -0.0054 Tc -0.0352 Tw (\), but their wild populations)Tj -9.5089 -1.0926 TD -0.0057 Tc -0.0301 Tw (are also in jeopardy \(Hill & Robertson, 1988\). More)Tj 0 -1.09 TD -0.0032 Tc 0.1227 Tw [(recent work, in particular by the British )9.7(T)35.6(rust for)]TJ T* 0.0077 Tc [(Ornithology)56.7(, has revealed further decreases \(e.g.)]TJ 0 -1.0926 TD -0.0061 Tc 0.021 Tw [(Chamberlain & Fuller)35.1(, 2000; Gregory )]TJ /F11 1 Tf 15.2968 0 TD 0.0227 Tw (et al.)Tj /F9 1 Tf 1.9894 0 TD -0.0052 Tc (, 2000\).)Tj -17.2862 -1.09 TD 0.0299 Tc 0.1227 Tw (Such population declines parallel what is also)Tj T* 0.0164 Tc [(happening to many other)34.4(, often less thoroughly)]TJ 0 -1.0926 TD -0.0059 Tc -0.0221 Tw [(studied, birds associated with farmland. In all cases,)]TJ 0 -1.0901 TD 0.0382 Tc 0.1227 Tw (the reason has been clearly related to loss of)Tj T* 0.0232 Tc [(biodiversity)59.3(, attributed to the intensification of)]TJ 0 -1.0926 TD 0.0033 Tc (farming practices in recent decades \(Potts, 1997;)Tj 0 -1.09 TD -0.0067 Tc [(W)45(ilson )]TJ /F11 1 Tf 3.0769 0 TD -0.0061 Tc 0.0227 Tw (et al.)Tj /F9 1 Tf 1.992 0 TD -0.0056 Tc 0.0245 Tw (, 1999\). Increased herbicide usage, the)Tj -5.0689 -1.09 TD -0.0126 Tc -0.0611 Tw [(summer use of foliar insecticides, and a loss of under)8(-)]TJ 0 -1.0926 TD 0.0512 Tc 0.1228 Tw (sowing, have all been implicated in causing)Tj 0 -1.09 TD -0.0062 Tc 0.116 Tw [(increased game chick mortality)55.6(, by decreasing the)]TJ T* 0.0077 Tc 0.1227 Tw (available invertebrate food \(Rands, 1986, 1988\).)Tj 22.422 29.452 TD -0.0058 Tc 0.0444 Tw (Grey partridge and pheasant \()Tj /F11 1 Tf 11.8952 0 TD 0.0455 Tw (Phasianus colchicus)Tj /F9 1 Tf 8.1329 0 TD 0 Tc 0 Tw (\))Tj -20.0281 -1.09 TD -0.0056 Tc 0.0233 Tw (chicks depend upon a high number of invertebrates)Tj T* -0.005 Tw (in their diet in the first few weeks of life, to provide)Tj T* 0.0006 Tc 0.1228 Tw [(suf)13.5(ficient protein for survival \(Hill, 1985; Green,)]TJ 0 -1.0926 TD -0.0056 Tc 0.1185 Tw [(1984; Potts, 1986; )12.8(W)46.1(ilson )]TJ /F11 1 Tf 10.9727 0 TD -0.0062 Tc 0.1182 Tw (et al.)Tj /F9 1 Tf 2.0873 0 TD -0.0053 Tc 0.1197 Tw (, 1999\). For other)Tj -13.06 -1.09 TD -0.0058 Tc 0.0916 Tw (species such as the red-legged partridge \()Tj /F11 1 Tf 16.8017 0 TD -0.0072 Tc (Alectoris)Tj -16.8017 -1.0901 TD -0.0069 Tc (rufa)Tj /F9 1 Tf 1.6467 0 TD -0.0059 Tc 0.0551 Tw (\), insects are also important, though to a lesser)Tj -1.6467 -1.0926 TD -0.0057 Tc -0.0091 Tw [(extent \(Green, 1984; Rands, 1988\). )12.3(The presence of)]TJ 0 -1.0901 TD 0.0047 Tc 0.1227 Tw (weeds in and around crops as hosts for non-pest)Tj T* -0.0061 Tc 0.0907 Tw [(insects is thus important. )47.8(Additionally)58.3(, the amount)]TJ 0 -1.0926 TD 0.1227 Tw (of plant material eaten increases as chicks mature)Tj 0 -1.09 TD 0.0331 Tc (\(Ford )Tj /F11 1 Tf 2.796 0 TD 0.0316 Tc (et al.)Tj /F9 1 Tf 2.3218 0 TD 0.0332 Tc (, 1938\), including grass and small)Tj -5.1178 -1.09 TD -0.012 Tc -0.0614 Tw (dicotyledonous seeds, unripe cereal grains and leaves)Tj 0 -1.0926 TD -0.006 Tc 0.1033 Tw [(and flowers \(Green, 1984\). )48.2(Adult game birds feed)]TJ 0 -1.09 TD 0.0158 Tc 0.1227 Tw (almost exclusively on plant material, with little)Tj T* -0.0068 Tc -0.0608 Tw [(dif)11.2(ference in preference between species \(Middleton)]TJ 0 -1.0926 TD -0.006 Tc 0.038 Tw [(& Chitty)55.8(, 1937\). Diet items consist of grain, leaves)]TJ 0 -1.09 TD -0.0056 Tc 0.0492 Tw (and roots of grasses and dicotyledonous plants.)Tj 0.8246 -1.0901 TD 0.1162 Tw (Later in the season, partridges pair for breeding)Tj -0.8246 -1.0926 TD 0.0937 Tw (and establish territories, so that the area of habitat)Tj 0 -1.0901 TD 0.0392 Tc 0.1227 Tw (accessible for nesting and brood rearing may)Tj T* 0.0226 Tc (constrain maximum population growth \(Rands,)Tj 0 -1.0926 TD -0.006 Tc 0.0416 Tw [(1986; )48(Aebischer & Blake, 1994\). Breeding success)]TJ 0 -1.09 TD 0.0254 Tw (during the spring-summer has been shown to relate)Tj T* 0.0042 Tc 0.1227 Tw (to the availability of nesting cover in this spring)Tj 0 -1.0926 TD 0.0105 Tc (settling period, as well as the likelihood of nest)Tj 0 -1.09 TD -0.0105 Tc -0.0609 Tw [(predation \(Potts, 1997\). )12.7(The structural characteristics)]TJ T* 0.0088 Tc 0.1227 Tw (of hedgerows are important in determining their)Tj ET endstream endobj 3 0 obj << /ProcSet [/PDF /Text ] /Font << /F4 4 0 R /F5 5 0 R /F6 6 0 R /F7 7 0 R /F8 8 0 R /F9 9 0 R /F10 10 0 R /F11 11 0 R /F12 12 0 R >> /ExtGState << /GS2 13 0 R >> >> endobj 16 0 obj << /Length 11810 >> stream BT /F4 1 Tf 9 0 0 9 59.502 788.95 Tm 0 0 0 1 k /GS2 gs -0.0019 Tc (112)Tj /F6 1 Tf 8 0 0 8 258.948 788.95 Tm -0.0084 Tc 0.0207 Tw [(S R )11.3(THOMAS )]TJ /F5 1 Tf 6.1618 0 TD -0.0051 Tc 0.0052 Tw [(E)5.3(T)12.6( AL.)]TJ /F9 1 Tf 11 0 0 11 59.502 762.985 Tm -0.0025 Tc 0.1227 Tw (suitability for nesting. Hedge-bottoms, with some)Tj 0 -1.09 TD 0.056 Tc 0.1728 Tw (elevation for good drainage, and with high)Tj T* -0.0075 Tc -0.061 Tw (proportions of dead grass, litter and other vegetation,)Tj 0 -1.0926 TD -0.0053 Tc 0.1227 Tw (have been found to be important, as birds require)Tj 0 -1.09 TD -0.0061 Tc 0.0058 Tw (shelter and material to conceal nests from predators)Tj T* -0.0057 Tc -0.0559 Tw [(\(Rands, 1986\). )48.4(Aebischer & Blake \(1994\) found that)]TJ 0 -1.0926 TD -0.006 Tc -0.0396 Tw [(several varieties of field mar)12(gins and non-crop areas)]TJ 0 -1.0901 TD 0.0554 Tc 0.2418 Tw (were preferred nesting habitat. Nests were)Tj T* 0.1354 Tw (particularly associated with species such as)Tj 0 -1.0926 TD 0.0157 Tc 0.123 Tw [(cock\222)46.6(sfoot \()]TJ /F11 1 Tf 5.074 0 TD 0.0164 Tc 0.1227 Tw (Dactylis glomerata)Tj /F9 1 Tf 8.1174 0 TD 0.0167 Tc 0.1228 Tw (\), nettles \()Tj /F11 1 Tf 4.4762 0 TD 0.0154 Tc [(U)7.8(r)-7.8(tica)]TJ -17.6676 -1.09 TD 0.0061 Tc (dioica)Tj /F9 1 Tf 2.546 0 TD 0.0079 Tc 0.1227 Tw (\) and hedge-parsley \()Tj /F11 1 Tf 9.0039 0 TD 0.0077 Tc 0.1228 Tw [(T)90.2(orilis japonica)]TJ /F9 1 Tf 6.5815 0 TD (\), i.e.)Tj -18.1314 -1.09 TD -0.006 Tc -0.0143 Tw [(those with tall, more continuous canopy cover)48.1(. Less)]TJ 0 -1.0926 TD -0.0079 Tc -0.061 Tw [(preferred vegetation was more open, low and patchy)56.5(.)]TJ 0 -1.0901 TD 0.0553 Tc 0.3114 Tw (Beetle banks may also fulfil such habitat)Tj T* -0.0059 Tc 0.0477 Tw [(requirements. Grassy banks and hedgerows are not)]TJ 0 -1.0926 TD 0.0871 Tw (such preferred habitats for pheasants, which select)Tj 0 -1.09 TD 0.0031 Tc 0.1226 Tw [(field mar)10.8(gins specifically adjacent to woodlands.)]TJ T* 0.0215 Tc 0.1229 Tw [(However)37(, these game birds do use this type of)]TJ 0 -1.0926 TD -0.006 Tc 0.0065 Tw [(vegetation where grass is suf)12(ficiently tall and dense)]TJ 0 -1.09 TD 0.0457 Tc 0.1227 Tw [(\(Hill & Robertson, 1988\). )9.6(V)112.8(egetation is also)]TJ T* -0.0053 Tc 0.1228 Tw [(important for chicks, as it must provide suf)10.2(ficient)]TJ 0 -1.0926 TD -0.001 Tc 0.1227 Tw (cover to conceal them from predators, yet permit)Tj 0 -1.09 TD 0.0146 Tc (movement of the chicks within it \(Aebischer &)Tj T* -0.0058 Tc 0.0682 Tw (Blake, 1994\).)Tj 0.822 -1.09 TD -0.0061 Tc 0.1227 Tw (More non-crop habitat and sympathetic farming)Tj -0.822 -1.0926 TD -0.0058 Tc -0.0409 Tw (practice is needed for the successful survival of wild)Tj 0 -1.0901 TD -0.0009 Tc 0.1226 Tw [(game species in the arable environment. )9.3(The UK)]TJ T* -0.0075 Tc -0.0613 Tw [(Ministry of )46.7(Agriculture, Fisheries and Food \(MAFF\))]TJ 0 -1.0926 TD -0.0062 Tc -0.0309 Tw [(is currently piloting an )48.1(Arable S)11.8(tewardship Scheme,)]TJ 0 -1.09 TD 0.0354 Tc 0.1227 Tw (encouraging farmers to manage their land for)Tj T* 0.0174 Tc (wildlife including game, by recommending and)Tj 0 -1.0926 TD -0.0045 Tc (monitoring a variety of approaches, such as grass)Tj 0 -1.09 TD -0.0025 Tc [(m)6.7(a)6.7(r)10.4(gins, wildlife seed mixtures for mar)10.4(gin strips,)]TJ T* 0.0032 Tc 0.1228 Tw (overwintered stubbles, undersown spring cereals,)Tj 0 -1.0926 TD 0.0375 Tc 0.1227 Tw (conservation headlands and also beetle banks)Tj 0 -1.09 TD 0.0558 Tc 0.1455 Tw (\(Anon., 1999)Tj /F11 1 Tf 6.128 0 TD 0 Tc 0 Tw (a)Tj /F9 1 Tf 0.554 0 TD 0.0556 Tc 0.1499 Tw [(\). )14.3(The benefits of many such)]TJ -6.682 -1.09 TD 0.0207 Tc 0.1225 Tw [(techniques, particularly of the latter)36.2(, are as yet)]TJ 0 -1.0926 TD -0.0041 Tc 0.1228 Tw (inadequately quantified.)Tj /F11 1 Tf 7.6329 -2.1801 TD -0.0065 Tc 0.0863 Tw (Beetle banks)Tj /F9 1 Tf -6.8109 -1.0926 TD -0.0059 Tc 0.1121 Tw (Beetle banks are grass-sown ridges, designed to)Tj -0.822 -1.09 TD -0.0056 Tc -0.03 Tw (provide additional overwintering sites for beneficial)Tj T* 0.0106 Tc 0.1227 Tw (predatory arthropods; and through their location)Tj 0 -1.0926 TD -0.0059 Tc -0.0494 Tw (across the centre of fields, reduce the distances these)Tj 0 -1.09 TD 0.043 Tc 0.1228 Tw (arthropods must disperse to achieve an early)Tj T* 0.0336 Tc 0.1227 Tw (homogeneous distribution within arable fields)Tj 0 -1.0926 TD -0.0079 Tc (\(Thomas )Tj /F11 1 Tf 3.7469 0 TD -0.0065 Tc -0.0091 Tw (et al.)Tj /F9 1 Tf 1.9585 0 TD -0.0054 Tc -0.0073 Tw (, 1991\). In the UK, the total length of)Tj -5.7054 -1.0901 TD 0.014 Tc 0.1228 Tw (hedgerows has severely decreased \(Bannister &)Tj T* -0.0047 Tc 0.1227 Tw [(W)83.1(att, 1994\), with many of those remaining either)]TJ 0 -1.0926 TD -0.0059 Tc -0.0319 Tw (lacking bottom vegetation or overwhelmed by weed)Tj 0 -1.09 TD 0.0148 Tc 0.1227 Tw [(species that threaten the crop or are dif)9.6(ficult to)]TJ T* -0.0109 Tc -0.0614 Tw (control, because of poor management \(Pollard, 1968;)Tj 0 -1.0926 TD 0.0406 Tc 0.1228 Tw [(Dunkley)56.1(, 1997\). Beetle banks compensate by)]TJ 0 -1.09 TD 0.0021 Tc (providing herbaceous and perennial hedgebottom)Tj T* -0.0059 Tc -0.0568 Tw (vegetation in a simple, inexpensive form \(Sotherton,)Tj 0 -1.0926 TD 0.0116 Tc 0.1227 Tw (1995\). Crop pest predators, such as carabid and)Tj 0 -1.0901 TD -0.0057 Tc 0.0674 Tw (staphylinid beetles and spiders, shelter over winter)Tj T* -0.0443 Tw (in low vegetation and survive best in dense tussocky)Tj 22.422 62.1792 TD 0.0254 Tc 0.123 Tw [(grass \(Luf)12.5(f, 1965; )48.9(Asteraki )]TJ /F11 1 Tf 12.2276 0 TD 0.1227 Tw (et al.)Tj /F9 1 Tf 2.2832 0 TD 0.0267 Tc 0.1228 Tw (, 1992, 1995;)Tj -14.5108 -1.09 TD -0.0083 Tc (Thomas )Tj /F11 1 Tf 3.3655 0 TD -0.0065 Tc -0.0591 Tw [(et al)]TJ /F9 1 Tf 1.6621 0 TD -0.0061 Tc -0.061 Tw [(. 1991, 1992\). )17.5(T)37.8(ussock-forming species)]TJ -5.0276 -1.09 TD 0.0296 Tc 0.1226 Tw [(such as cock\222)45.1(sfoot, and )32.1(Y)99.3(orkshire fog, )]TJ /F11 1 Tf 17.379 0 TD 0.0288 Tc (Holcus)Tj -17.379 -1.0926 TD -0.0061 Tc (lanatus)Tj /F9 1 Tf 2.9094 0 TD 0.0621 Tw (, therefore, are recommended for sowing in)Tj -2.9094 -1.09 TD -0.0059 Tc -0.0514 Tw [(beetle banks \(Sotherton, 1995; )12.6(Thomas )]TJ /F11 1 Tf 15.4514 0 TD -0.007 Tc -0.0546 Tw (et al)Tj /F9 1 Tf 1.6673 0 TD -0.0052 Tc -0.0545 Tw (., 1991\).)Tj -17.1187 -1.09 TD -0.0057 Tc 0.0242 Tw (Previous studies on beetle banks have concentrated)Tj 0 -1.0926 TD -0.0059 Tc 0.1076 Tw (primarily on their functional value compared with)Tj 0 -1.0901 TD -0.0053 Tc 0.0617 Tw (other types of field boundaries for supporting high)Tj T* 0.0216 Tc 0.1227 Tw (densities of overwintering beneficial predators,)Tj 0 -1.0926 TD -0.0057 Tc 0.0436 Tw (although none have monitored invertebrates within)Tj 0 -1.09 TD -0.0117 Tc -0.0591 Tw (spring \(Thomas )Tj /F11 1 Tf 6.2697 0 TD -0.0121 Tc -0.0636 Tw (et al.)Tj /F9 1 Tf 1.8709 0 TD -0.011 Tc -0.0613 Tw (, 1991; MacLeod, 1994; Collins)Tj /F11 1 Tf -8.1406 -1.09 TD -0.0065 Tc 0.0091 Tw (et al.)Tj /F9 1 Tf 1.9791 0 TD -0.0057 Tc 0.0102 Tw (, 1996\). Other research to date has shown how)Tj -1.9791 -1.0926 TD -0.0058 Tc -0.037 Tw (beetle banks may provide nesting habitat for harvest)Tj 0 -1.0901 TD 0.0505 Tc 0.1227 Tw (mice \(Bence )Tj /F11 1 Tf 6.1331 0 TD 0.0517 Tc (et al.)Tj /F9 1 Tf 2.4404 0 TD 0.0528 Tc (, 1999\) and has indicated)Tj -8.5735 -1.09 TD 0.0555 Tc 0.4523 Tw (successional change in their vegetational)Tj 0 -1.0926 TD 0.0258 Tc 0.1227 Tw (composition \(Thomas )Tj /F11 1 Tf 9.7692 0 TD 0.0264 Tc (et al.)Tj /F9 1 Tf 2.2858 0 TD 0.1226 Tw [(, 2000\). )13.4(The Game)]TJ -12.055 -1.09 TD 0.0404 Tc 0.1228 Tw [(Conservancy )12.1(T)37.9(rust and regional Farming and)]TJ T* -0.0058 Tc 0.0752 Tw [(W)45.9(ildlife )48.5(Advisory Groups now propose that beetle)]TJ 0 -1.0926 TD 0.0281 Tc 0.1227 Tw (banks may be useful for gamebirds, providing)Tj 0 -1.09 TD 0.0174 Tc (additional nesting and feeding sites, if they are)Tj T* 0.0389 Tc 0.1228 Tw (retained as permanent landscape features and)Tj 0 -1.0926 TD 0.0487 Tc 0.1227 Tw [(protected from pesticide and herbicide drift.)]TJ 0 -1.09 TD -0.0061 Tc 0.0216 Tw [(Application of chemicals within them is restricted)]TJ T* 0.0142 Tc 0.1228 Tw (to the localised treatment of specific pernicious)Tj T* -0.0072 Tc -0.0608 Tw [(weeds. )10.9(The Countryside S)13.4(tewardship Scheme, along)]TJ 0 -1.0926 TD 0.0373 Tc 0.1227 Tw [(with the pilot )50.2(Arable S)14.1(tewardship Scheme, is)]TJ 0 -1.0901 TD 0.0003 Tc (currently alone in providing financial support for)Tj T* -0.0057 Tc 0.0565 Tw (setting up beetle banks on arable farmland \(Anon.,)Tj 0 -1.0926 TD -0.0061 Tc (1999)Tj /F11 1 Tf 1.9817 0 TD 0 Tc 0 Tw (a)Tj /F9 1 Tf 0.4974 0 TD -0.0062 Tc 0.0341 Tw [( and )]TJ /F11 1 Tf 1.9843 0 TD 0 Tc 0 Tw (b)Tj /F9 1 Tf 0.4973 0 TD -0.0104 Tc (\).)Tj /F11 1 Tf 2.5177 -2.1801 TD -0.0064 Tc 0.0432 Tw (Aims of study)Tj /F9 1 Tf -6.6537 -1.0926 TD -0.0057 Tc -0.015 Tw (This study was designed to quantify the resources)Tj -0.8246 -1.09 TD 0.0046 Tc 0.1229 Tw [(of)9.8(fered by beetle banks for gamebird chicks and)]TJ T* -0.0062 Tc 0.1227 Tw [(adults. Firstly)55.6(, it aimed to measure the abundance)]TJ 0 -1.0926 TD 0.0114 Tc (of chick food invertebrates in beetle banks; and)Tj 0 -1.09 TD -0.0121 Tc -0.0608 Tw [(secondly)60.1(, to quantify their vegetational value for both)]TJ T* -0.0103 Tc -0.0609 Tw [(chicks and adult birds. )13(The latter involved examining)]TJ 0 -1.0926 TD -0.006 Tc [(cover)32.7(, used for nesting and brood-rearing, and plants)]TJ 0 -1.09 TD 0.0557 Tc 0.3709 Tw (of food value. Relationships between the)Tj T* -0.0061 Tc -0.061 Tw (invertebrates and plant cover could then be assessed.)Tj 0 -1.0926 TD -0.0477 Tw (These attributes were compared with those provided)Tj 0 -1.09 TD -0.0097 Tc -0.0614 Tw [(in adjacent typical conventional field mar)13.5(gins, which)]TJ T* 0.0472 Tc 0.1227 Tw (are also linear vegetation strips with similar)Tj 0 -1.0926 TD -0.0057 Tc 0.0694 Tw (functions in the agricultural landscape.)Tj /F12 1 Tf 5.1951 -2.1801 TD -0.0068 Tc 0.0773 Tw (Materials and Methods)Tj /F11 1 Tf -3.2624 -2.1827 TD -0.0059 Tc 0.0682 Tw (Chick food availability in spring/summer)Tj /F9 1 Tf -1.1081 -1.09 TD 0.0194 Tc 0.1227 Tw (Sampling took place in beetle banks on farm)Tj -0.8246 -1.0926 TD -0.0063 Tc 0.0448 Tw [(estates across Hampshire and )9.2(W)48(iltshire, UK, all on)]TJ 0 -1.09 TD -0.0079 Tc -0.0613 Tw [(slightly flinty)59.1(, calcareous silty clay loam-based soils.)]TJ T* -0.006 Tc 0.023 Tw (Ranging from 200 m - 900 m long and 2.5 m - 5 m)Tj 0 -1.0926 TD -0.0059 Tc 0.0766 Tw (wide, the banks had been established between less)Tj 0 -1.09 TD -0.0053 Tc 0.1058 Tw (than 1 and 14 yr previously by autumn-ploughing)Tj T* 0.0554 Tc 0.4427 Tw [(earth ridges. )9(They were hand-sown with)]TJ 0 -1.0926 TD -0.0067 Tc (predominately)Tj /F11 1 Tf 5.6951 0 TD -0.0061 Tc -0.0523 Tw [( Dactylis glomerata)]TJ /F9 1 Tf 7.7283 0 TD -0.0056 Tc -0.0546 Tw [( and various other)]TJ -13.4233 -1.0901 TD -0.006 Tc 0.0606 Tw (grass species. Natural regeneration has taken place)Tj T* 0 Tc 0.1227 Tw (within them, although there has been little active)Tj ET endstream endobj 17 0 obj << /ProcSet [/PDF /Text ] /Font << /F4 4 0 R /F5 5 0 R /F6 6 0 R /F9 9 0 R /F11 11 0 R /F12 12 0 R >> /ExtGState << /GS2 13 0 R >> >> endobj 19 0 obj << /Length 13400 >> stream BT /F4 1 Tf 9 0 0 9 522.205 788.95 Tm 0 0 0 1 k /GS2 gs -0.0019 Tc (113)Tj /F5 1 Tf 8 0 0 8 230.038 788.95 Tm -0.0057 Tc 0.0954 Tw [(Resour)29.7(ce pr)29.7(ovision for farmland gamebir)33.3(d)8.3(s)]TJ /F9 1 Tf 11 0 0 11 311.789 762.985 Tm -0.0045 Tc 0.1227 Tw (placed on the ground was recorded. Overall plant)Tj 0 -1.09 TD -0.0059 Tc 0.117 Tw [(cover)32.8(, the amount of live/dead tussock, and cover)]TJ T* 0.0045 Tc 0.1227 Tw (of other grasses and dicotyledonous plants, were)Tj 0 -1.0926 TD 0.0088 Tc (compared between habitats by two-tailed )Tj /F11 1 Tf 17.6598 0 TD 0 Tc 0 Tw (t)Tj /F9 1 Tf 0.286 0 TD 0.0081 Tc (-tests.)Tj -17.9459 -1.09 TD -0.0035 Tc 0.1228 Tw (Mean values from each site were used, following)Tj T* -0.0057 Tc 0.1227 Tw (logit transformation.)Tj /F12 1 Tf 8.6199 -2.1827 TD 0.0012 Tc (Results)Tj /F11 1 Tf -6.9732 -2.1827 TD -0.0057 Tc 0.0537 Tw (Spring and summer chick food availability)Tj /F9 1 Tf -0.8221 -1.09 TD -0.0447 Tw (In 1998, chickfood invertebrate densities were not)Tj -0.8246 -1.09 TD 0.0051 Tc 0.1226 Tw [(significantly dif)15.4(ferent between field mar)7.7(gins and)]TJ 0 -1.0926 TD -0.0061 Tc 0.1159 Tw (beetle banks \(F)Tj /F10 1 Tf 6.4 0 0 6.4 380.586 615.895 Tm -0.0018 Tc 0.125 Tw (1, 6)Tj /F9 1 Tf 11 0 0 11 390.989 623.237 Tm 0 Tc 0 Tw ( )Tj 0.3633 -0.3865 TD -0.0059 Tc 0.1182 Tw (= 0.02, )Tj /F11 1 Tf 3.2598 0 TD 0 Tc 0 Tw (P)Tj /F9 1 Tf 0.6056 0 TD -0.0064 Tc 0.1182 Tw [( = 0.89\). )14.2(There was a)]TJ -11.4288 -1.0901 TD -0.0058 Tc 0.1 Tw (significantly higher prey density per m)Tj /F10 1 Tf 6.4 0 0 6.4 485.638 610.651 Tm 0 Tc 0 Tw (2)Tj /F9 1 Tf 11 0 0 11 488.841 606.994 Tm -0.0067 Tc 0.1001 Tw [( in )47.5(August)]TJ -16.0956 -1.09 TD 0.0015 Tw (compared to May \(F)Tj /F10 1 Tf 6.4 0 0 6.4 400.684 591.886 Tm -0.0015 Tc 0 Tw (1, 6)Tj /F9 1 Tf 11 0 0 11 410.293 595.004 Tm -0.0056 Tc 0.0013 Tw ( = 7.03, )Tj /F11 1 Tf 3.2727 0 TD 0 Tc 0 Tw (P)Tj /F9 1 Tf 0.6056 0 TD -0.0056 Tc 0.0021 Tw ( = 0.04\), caused by)Tj -12.8332 -1.0926 TD 0.0151 Tc 0.1227 Tw (increased in most taxa, although the interaction)Tj 0 -1.09 TD -0.0058 Tc 0.0894 Tw (between habitat and date was non-significant \(F)Tj /F10 1 Tf 6.4 0 0 6.4 525.493 567.905 Tm -0.0017 Tc 0.0937 Tw (1, 6)Tj /F9 1 Tf 11 0 0 11 311.789 559.004 Tm -0.006 Tc -0.0205 Tw (= 0.00, )Tj /F11 1 Tf 2.9815 0 TD 0 Tc 0 Tw (P)Tj /F9 1 Tf 0.6056 0 TD -0.0061 Tc -0.0214 Tw [( = 0.99\). In May)58.3(, mean chickfood densities)]TJ -3.5871 -1.0926 TD -0.0016 Tw (were 657.50 per m)Tj /F10 1 Tf 6.4 0 0 6.4 393.087 550.642 Tm 0 Tc 0 Tw (2)Tj /F9 1 Tf 11 0 0 11 396.29 546.985 Tm -0.0055 Tc -0.0021 Tw ( for beetle banks and 564.17 per)Tj -7.6819 -1.09 TD 0 Tc 0 Tw (m)Tj /F10 1 Tf 6.4 0 0 6.4 320.435 538.651 Tm (2)Tj /F9 1 Tf 11 0 0 11 323.837 534.994 Tm 0.0205 Tc 0.1227 Tw [( for field mar)12.8(gins \()]TJ /F13 1 Tf 8.8 0 0 8.8 416.756 534.994 Tm 0.0256 Tc (SED)Tj /F9 1 Tf 11 0 0 11 434.104 534.994 Tm 0.0211 Tc 0.1231 Tw [( = 232.7\). In )49.8(August,)]TJ -11.1195 -1.09 TD -0.0054 Tc 0.076 Tw (densities were 1547.31 and 1434.23 for banks and)Tj 0 -1.0926 TD -0.0059 Tc 0.0477 Tw [(m)7.1(a)7.1(r)12.1(gins, respectively \()]TJ /F13 1 Tf 8.8 0 0 8.8 411.597 510.985 Tm -0.0082 Tc (SED)Tj /F9 1 Tf 11 0 0 11 428.038 510.985 Tm -0.0057 Tc 0.0488 Tw [( = 450.2\). Prey densities)]TJ -10.5681 -1.09 TD -0.0059 Tc -0.025 Tw (showed high variability between the fields sampled,)Tj T* 0.04 Tw (on both of the sampling occasions.)Tj 0.8246 -1.09 TD -0.0033 Tc 0.1227 Tw (Small flies were very frequently caught, and as)Tj -0.8246 -1.0926 TD -0.0064 Tc -0.0552 Tw (might be expected, were predominant in the summer)Tj 0 -1.0901 TD -0.006 Tc -0.0481 Tw [(catch \(T)63.6(able 1\). Small species of staphylinid beetles,)]TJ T* -0.0055 Tc -0.0201 Tw (bugs and linyphiid spiders were also very abundant.)Tj 0 -1.0926 TD -0.0061 Tc 0.0755 Tw (Hymenopteran larvae, regarded as important chick)Tj 0 -1.09 TD -0.0063 Tc -0.0482 Tw [(food components \(Moreby & )48.1(Aebischer)32.4(, 1992\) were)]TJ T* -0.0128 Tc -0.061 Tw (infrequently caught, and were most numerous in field)Tj 0 -1.0926 TD 0.0308 Tc 0.1227 Tw [(m)7.8(a)7.8(r)12.8(gins in May)56.6(. Beetle banks contained more)]TJ 0 -1.09 TD -0.0093 Tc -0.061 Tw (carabid beetle and heteropteran bugs in both seasons,)Tj T* 0.0411 Tc 0.1227 Tw (and more coccinellid beetles in spring, when)Tj 0 -1.0926 TD 0.0103 Tc [(compared with the field mar)15.5(gins. Only ants and)]TJ 0 -1.09 TD -0.0093 Tc -0.0609 Tw (lepidopteran larvae were consistently more abundant)Tj T* -0.0067 Tc 0.0499 Tw [(in field mar)8.8(gins.)]TJ 0.8246 -1.0926 TD -0.0057 Tc 0.0597 Tw [(In 1999, sweep capture of mean total chick-food)]TJ -0.8246 -1.09 TD -0.0059 Tc 0.0264 Tw (invertebrates was significantly higher in permanent)Tj T* -0.0064 Tc 0.1221 Tw [(field mar)11.6(gins compared with beetle banks \(F)]TJ /F10 1 Tf 6.4 0 0 6.4 513.503 291.895 Tm -0.0022 Tc 0.125 Tw (1, 34 )Tj /F12 1 Tf 11 0 0 11 529.49 295.013 Tm 0 Tc 0 Tw (=)Tj /F9 1 Tf -19.791 -1.0926 TD 0.0481 Tc (7.20, )Tj /F11 1 Tf 2.6697 0 TD 0 Tc (P)Tj /F9 1 Tf 0.6571 0 TD 0.0488 Tc 0.1228 Tw [( = 0.01\). )48.9(A)56.7( mean of 46.65 chickfood)]TJ -3.3268 -1.09 TD -0.0016 Tc 0.1227 Tw (invertebrates was caught per 15 samples taken in)Tj T* -0.006 Tc -0.0262 Tw [(beetle banks, with a mean for field mar)12(gins of 64.70)]TJ 0 -1.0926 TD 0 Tc 0 Tw (\()Tj /F13 1 Tf 8.8 0 0 8.8 315.389 246.995 Tm -0.0082 Tc (SED)Tj /F9 1 Tf 11 0 0 11 331.859 246.995 Tm -0.006 Tc -0.0286 Tw [( = 8.73\). Catch did not dif)12(fer between the farms)]TJ -1.8245 -1.09 TD -0.0076 Tc 0.0727 Tw (sampled \(F)Tj /F10 1 Tf 6.4 0 0 6.4 361.055 231.914 Tm -0.0021 Tc 0.0703 Tw (4, 34)Tj 2.0684 -0.6644 TD 0 Tc 0 Tw ( )Tj /F12 1 Tf 11 0 0 11 376.391 235.004 Tm -0.0107 Tc (= )Tj /F9 1 Tf 0.8813 0 TD -0.0052 Tc (2.30, )Tj /F11 1 Tf 2.3012 0 TD 0 Tc (P)Tj /F9 1 Tf 0.603 0 TD -0.0057 Tc 0.0742 Tw [( = 0.08\), and there was no)]TJ -9.6584 -1.0901 TD 0.0168 Tc 0.0873 Tw (significant interaction between habitat and farm)Tj 0 -1.0926 TD -0.013 Tc (\(F)Tj /F10 1 Tf 6.4 0 0 6.4 321.484 207.905 Tm 0.0034 Tc 0.125 Tw (4, 634)Tj 2.6575 -0.6644 TD 0 Tc 0 Tw ( )Tj /F12 1 Tf 11 0 0 11 340.901 210.995 Tm -0.0066 Tc (= )Tj /F9 1 Tf 0.9406 0 TD -0.0009 Tc (0.59, )Tj /F11 1 Tf 2.3734 0 TD 0 Tc (P)Tj /F9 1 Tf 0.6082 0 TD -0.0017 Tc 0.1227 Tw [( = 0.67\). )52.4(As in the first sampling)]TJ -6.5686 -1.0901 TD 0.0108 Tc 0.1229 Tw [(year)31.4(,)-0.2( there was considerable variability between)]TJ T* -0.0069 Tc 0.0909 Tw (sampling sites.)Tj 0.8246 -1.0926 TD -0.006 Tc -0.0325 Tw (Small species of diptera, heteropterans and aphids)Tj -0.8246 -1.09 TD -0.0064 Tc 0.0994 Tw [(were most numerous in the sweepnet catch \(T)60.6(able)]TJ T* -0.0067 Tc -0.0611 Tw [(2\). Beetles were frequently caught, and did not dif)13.9(fer)]TJ 0 -1.0926 TD -0.0048 Tc 0.1229 Tw [(in abundance between habitat type. Field mar)10.7(gins)]TJ 0 -1.09 TD -0.0058 Tc -0.0509 Tw (contained significantly greater numbers of cantharid)Tj T* 0.0254 Tc 0.1227 Tw (beetles, heteropterans, other auchenorrhynchan)Tj 0 -1.0926 TD -0.0063 Tc 0.0573 Tw [(bugs, flies and some spider families. )11.7(There was no)]TJ 0 -1.0901 TD -0.0056 Tc 0.0728 Tw [(dif)12.4(ference in the abundance of infrequently caught)]TJ T* 0.0257 Tc 0.1227 Tw (sawfly or lepidopteran larvae between the two)Tj -22.422 62.1792 TD -0.0126 Tc -0.0613 Tw (management. In all cases, adjacent established grassy)Tj 0 -1.09 TD -0.006 Tc 0.1119 Tw [(hedgerow bottoms or grassy non-shrubby mar)12(gins)]TJ T* -0.0064 Tc 0.0809 Tw (were sampled simultaneously to allow comparison)Tj 0 -1.0926 TD -0.0059 Tc 0.047 Tw (with the beetle banks.)Tj 0.822 -1.09 TD -0.0061 Tc 0.0144 Tw [(In 1998, four beetle banks/mar)11.9(gins were sampled)]TJ -0.822 -1.09 TD 0.0149 Tc 0.1226 Tw [(in May and five in )45.7(August, to assess chickfood)]TJ 0 -1.0926 TD -0.0055 Tc 0.1027 Tw (provision through the main chick-hatching period,)Tj 0 -1.0901 TD -0.0062 Tc 0.1147 Tw (on a single Hampshire estate. Fifteen 20 cm \327 20)Tj T* -0.0063 Tc -0.012 Tw [(cm areas along each beetle bank or field mar)11.7(gin site)]TJ 0 -1.0926 TD 0.0421 Tc 0.1227 Tw (were randomly selected. Invertebrates on the)Tj 0 -1.09 TD -0.0077 Tc -0.0613 Tw [(vegetation and soil surface were removed by a R)36.1(yobi)]TJ T* 0.0303 Tc 0.1227 Tw [(RSV3100 vacuum suction-sampler \(S)12.3(tewart &)]TJ 0 -1.0926 TD -0.0056 Tc 0.1006 Tw [(W)46.1(right, 1995\). Samples were frozen prior to hand)]TJ 0 -1.0901 TD -0.0059 Tc 0.0583 Tw (sorting to remove invertebrates, which were stored)Tj T* -0.0088 Tc -0.0613 Tw [(in 70% alcohol before identification. )50.5(All invertebrate)]TJ 0 -1.0926 TD -0.0006 Tc 0.1227 Tw (taxa on which game chicks most commonly feed)Tj 0 -1.09 TD -0.0059 Tc 0.0355 Tw (were identified predominantly to family \(following)Tj T* -0.0098 Tc (Ford )Tj /F11 1 Tf 2.0409 0 TD -0.0089 Tc -0.059 Tw (et al.)Tj /F9 1 Tf 1.8915 0 TD -0.0613 Tw [(, 1938; Green, 1984; Moreby & Southway)60.7(,)]TJ -3.9324 -1.0926 TD -0.006 Tc 0.0341 Tw (1999; Moreby )Tj /F11 1 Tf 5.9399 0 TD -0.0064 Tc 0.0318 Tw (et al.)Tj /F9 1 Tf 2.0023 0 TD -0.0059 Tc 0.0345 Tw (, 1999\). Mean total numbers of)Tj -7.9422 -1.09 TD 0.0421 Tc 0.1227 Tw (these chickfood invertebrates were compared)Tj T* -0.006 Tc -0.0022 Tw [(between beetle banks and field mar)12(gins, for the two)]TJ 0 -1.0926 TD -0.0089 Tc -0.0604 Tw [(sampling periods, using repeated measures )45.7(ANOV)122.5(A,)]TJ 0 -1.09 TD 0.0257 Tc 0.1227 Tw (following log \(x+1\) transformation to increase)Tj T* -0.006 Tc 0.0614 Tw (homogeneity of variance.)Tj 0.822 -1.09 TD -0.0533 Tw [(In 1999, 22 beetle banks/mar)12(gins from five estates)]TJ -0.822 -1.0926 TD -0.0007 Tc 0.1228 Tw (were assessed, to extend the data set and include)Tj 0 -1.0901 TD -0.0061 Tc -0.0611 Tw [(farm variation as a factor)48(. Sweep-netting was carried)]TJ T* 0.0149 Tc 0.1227 Tw (out over June and July to facilitate rapid insect)Tj 0 -1.0926 TD -0.006 Tc 0.022 Tw [(collection from a lar)12(ge number of sites during peak)]TJ 0 -1.09 TD 0.032 Tc 0.1225 Tw [(chick hatch. )44.7(A)52.8( 50 cm diameter net was swept)]TJ T* -0.0082 Tc -0.0614 Tw (immediately above the ground within the vegetation,)Tj 0 -1.0926 TD -0.0059 Tc 0.0455 Tw (taking approximately 1 s to collect from an area of)Tj 0 -1.09 TD 0.0048 Tc 0.1227 Tw (around 50 cm)Tj /F10 1 Tf 6.4 0 0 6.4 129.011 382.661 Tm 0 Tc 0 Tw (2)Tj /F9 1 Tf 11 0 0 11 132.243 379.004 Tm 0.0049 Tc 0.1227 Tw [( each sweep. Fifteen sweeps were)]TJ -6.0996 -1.09 TD -0.0137 Tc -0.0611 Tw (pooled to form a sample, with 15 samples being taken)Tj 0 -1.0926 TD -0.0086 Tc -0.061 Tw (randomly along each site. Samples were again stored)Tj 0 -1.09 TD -0.0058 Tc 0.0402 Tw (frozen and hand-sorted to remove all invertebrates,)Tj T* 0.0024 Tc 0.1227 Tw [(which were identified as previously)59.1(. )10.1(The relative)]TJ 0 -1.0926 TD -0.0056 Tc 0.0257 Tw (abundance of invertebrate prey available for chicks)Tj 0 -1.09 TD -0.006 Tc 0.0188 Tw (to feed on was compared between beetle banks and)Tj T* -0.0062 Tc 0.0091 Tw [(field mar)11.8(gins, and between dif)11.8(ferent farms, by two-)]TJ 0 -1.0926 TD -0.0064 Tc -0.0031 Tw [(way )48.4(ANOV)122.4(A)55.6( following log \(x+1\) transformation of)]TJ 0 -1.09 TD 0.027 Tc 0.1227 Tw (the data. Site means were again used to avoid)Tj T* 0.0066 Tc 0.0749 Tw [(pseudoreplication. Additionally)55.6(, )-47.7(Shannon-W)32.4(iener)]TJ 0 -1.0926 TD -0.0053 Tc 0.1227 Tw (Diversity indices were calculated for each sample)Tj 0 -1.09 TD 0.0083 Tc (of chickfood invertebrates, and used to compare)Tj T* -0.0068 Tc 0.0916 Tw [(between habitats and farms, by two-way )48(ANOV)119.5(A.)]TJ 0 -1.0926 TD -0.006 Tc 0.0439 Tw (Being normally distributed when calculated from a)Tj 0 -1.0901 TD -0.0059 Tc 0.0023 Tw (number of samples, this index does not require data)Tj T* 0.1022 Tw (transformation \(Magurran, 1988\).)Tj /F11 1 Tf 1.255 -2.1827 TD -0.0056 Tc 0.047 Tw [(V)110.5(egetation cover and food plant pr)30.5(ovision in)]TJ 6.4321 -1.09 TD -0.0063 Tc 0.0864 Tw (beetle banks)Tj /F9 1 Tf -6.865 -1.0926 TD 0.0327 Tc 0.1227 Tw (The vegetation within 20 randomly selected)Tj -0.822 -1.09 TD -0.0057 Tc 0.0108 Tw (sampling points on nine beetle banks and nine field)Tj T* -0.0375 Tw [(m)7.3(a)7.3(r)12.3(gins was assessed in July 1998, and in February/)]TJ 0 -1.0926 TD -0.0059 Tc -0.0101 Tw [(March 1999. )12.3(This was repeated for 22 beetle banks/)]TJ 0 -1.0901 TD -0.0057 Tc -0.0455 Tw [(m)7.3(a)7.3(r)12.3(gins in late June/July 1999. Each species present)]TJ T* -0.0058 Tc 0.109 Tw (and its percentage cover within a 0.71 m)Tj /F10 1 Tf 6.4 0 0 6.4 249.399 82.642 Tm 0 Tc 0 Tw (2)Tj /F9 1 Tf 11 0 0 11 252.602 79.013 Tm -0.0061 Tc 0.1091 Tw [( quadrat)]TJ ET endstream endobj 20 0 obj << /ProcSet [/PDF /Text ] /Font << /F4 4 0 R /F5 5 0 R /F9 9 0 R /F10 10 0 R /F11 11 0 R /F12 12 0 R /F13 21 0 R >> /ExtGState << /GS2 13 0 R >> >> endobj 23 0 obj << /Length 13938 >> stream BT /F4 1 Tf 9 0 0 9 59.502 788.95 Tm 0 0 0 1 k /GS2 gs -0.0019 Tc (114)Tj /F6 1 Tf 8 0 0 8 258.948 788.95 Tm -0.0084 Tc 0.0207 Tw [(S R )11.3(THOMAS )]TJ /F5 1 Tf 6.1618 0 TD -0.0051 Tc 0.0052 Tw [(E)5.3(T)12.6( AL.)]TJ /F14 1 Tf 10 0 0 10 71.918 763.637 Tm -0.0045 Tc 0.022 Tw [(T)92(able 1. Mean chickfood invertebrate densities per m)]TJ /F15 1 Tf 5.8 0 0 5.8 284.516 767.01 Tm 0 Tc 0 Tw (2)Tj /F14 1 Tf 10 0 0 10 287.408 763.637 Tm -0.0047 Tc 0.0233 Tw [( in mar)32.2(gin habitats sampled by R)15.1(yobi suction sampling in)]TJ -21.7843 -1.0998 TD 0.0142 Tw [(May and )18(August 1998. Results of )]TJ /F16 1 Tf 13.4192 0 TD 0 Tc 0 Tw (t)Tj /F14 1 Tf 0.275 0 TD -0.0044 Tc 0.0125 Tw (-test on log)Tj /F15 1 Tf 5.8 0 0 5.8 251.351 750.257 Tm -0.0049 Tc (10)Tj /F14 1 Tf 10 0 0 10 257.162 752.638 Tm -0.0044 Tc 0.015 Tw [( \(x + 1\) transformed data. \(*)]TJ /F16 1 Tf 11.52 0 TD 0 Tc 0 Tw (P)Tj /F14 1 Tf 0.5499 0 TD -0.0036 Tc 0.0133 Tw [( < 0.05, **)]TJ /F16 1 Tf 4.4334 0 TD 0 Tc 0 Tw (P)Tj /F14 1 Tf 0.5499 0 TD -0.004 Tc 0.015 Tw [( < 0.01, ***)]TJ /F16 1 Tf 4.9351 0 TD 0 Tc 0 Tw (P)Tj /F14 1 Tf 0.5499 0 TD -0.0057 Tc 0.015 Tw [( < 0.001\))]TJ ET 0 0 0 1 K 0 J 0 j 0.51 w 10 M []0 d /GS1 gs 1 i 59.502 741.243 m 530.053 741.243 l 59.502 727.098 m 530.053 727.098 l 59.502 457.495 m 530.053 457.495 l S BT /F2 1 Tf 9.012 0 0 9.012 242.507 370.783 Tm 0 g 0 Tc 0.0016 Tw (bb mean \()Tj 7 0 0 7 278.563 370.783 Tm -0.0012 Tc (SE)Tj 9.012 0 0 9.012 286.727 370.783 Tm -0.0004 Tc 0.002 Tw [(\))-9609.5(f)-0.8(m mean \()]TJ 7 0 0 7 413.379 370.783 Tm -0.0012 Tc (SE)Tj 9.012 0 0 9.012 421.543 370.783 Tm 0 Tc 0 Tw (\))Tj -39.7306 -1.8148 TD 0.0001 Tc [(Carabidae)-15367(0)0(.68)-2305.5(\()-0.3(0.22\))-8486.1(1)0(.22)-2305.5(\()-0.3(0.42\))-4806.2(t)]TJ 6.008 0 0 6.008 477.47 352.925 Tm (35)Tj 9.012 0 0 9.012 483.508 354.428 Tm -0.0009 Tc 0.0009 Tw [( = 2.03 n.)-5.7(s.)]TJ -46.6061 -1.4814 TD 0 Tc [(Staphylinidae)-13923.4(0.59)-2305.6(\(0.17\))-8486.2(1.06)-2305.6(\(0.30\))-4806.3(t)]TJ 6.008 0 0 6.008 477.47 339.574 Tm 0.0001 Tc (36)Tj 9.012 0 0 9.012 483.508 341.076 Tm -0.0009 Tc [( = 2.03 n.)-5.7(s.)]TJ -46.6061 -1.4814 TD -0.0002 Tc [(Chrysomelidae)-12870(13.16)-2308.9(\(8.87\))-7983.2(10.36)-2308.9(\()-0.6(4.14\))-4806.5(t)]TJ 6.008 0 0 6.008 477.47 326.223 Tm 0.0001 Tc (42)Tj 9.012 0 0 9.012 483.508 327.725 Tm -0.0009 Tc [( = 2.02 n.)-5.7(s.)]TJ -46.6061 -1.4783 TD -0.0002 Tc [(Curcurlionidae)-12926.6(12.33)-2308.9(\(7.36\))-8486.4(7.58)-2305.8(\(3.37\))-4806.5(t)]TJ 6.008 0 0 6.008 477.47 312.9 Tm 0.0001 Tc (35)Tj 9.012 0 0 9.012 483.508 314.402 Tm -0.0009 Tc [( = 2.03 n.)-5.7(s.)]TJ -46.6061 -1.4814 TD 0 Tc [(Cantharidae)-14590.2(2)-0.1(.84)-2305.6(\()-0.4(1.26\))-8486.2(6)-0.1(.66)-2305.6(\()-0.4(1.54\))-4806.3(t)]TJ 6.008 0 0 6.008 477.47 299.549 Tm 0.0001 Tc (42)Tj 9.012 0 0 9.012 483.508 301.051 Tm -0.0002 Tc 0.0002 Tw ( = 2.02 *)Tj -46.6061 -1.4783 TD 0 Tc [(Elateridae)-15370.3(1.65)-2305.6(\(0.47\))-8486.2(1.50)-2305.6(\(0.41\))-4806.3(t)]TJ 6.008 0 0 6.008 477.47 286.226 Tm 0.0001 Tc (42)Tj 9.012 0 0 9.012 483.508 287.728 Tm -0.0009 Tc 0.0009 Tw [( = 2.02 n.)-5.7(s.)]TJ -46.6061 -1.4814 TD -0.0001 Tc [(Coccinellidae)-13926.7(1.79)-2305.7(\(0.47\))-8486.3(1.70)-2305.7(\(0.70\))-4806.4(t)]TJ 6.008 0 0 6.008 477.47 272.875 Tm 0.0001 Tc (42)Tj 9.012 0 0 9.012 483.508 274.377 Tm -0.0009 Tc [( = 2.02 n.)-5.7(s.)]TJ -46.6061 -1.4814 TD -0.0001 Tc [(Nitidulidae)-14423.6(19.72)-1808.7(\(10.21\))-7983.1(36.41)-1805.6(\()-0.5(12.59\))-4806.4(t)]TJ 6.008 0 0 6.008 477.47 259.524 Tm 0.0001 Tc (42)Tj 9.012 0 0 9.012 483.508 261.026 Tm -0.0009 Tc [( = 2.02 n.)-5.7(s.)]TJ -46.6061 -1.4814 TD -0.0001 Tc [(Heteroptera)-14203.5(78.51)-1808.7(\(13.33\))-7486.2(112.05)-1802.4(\(12.59\))-4806.4(t)]TJ 6.008 0 0 6.008 477.47 246.173 Tm 0.0001 Tc (37)Tj 9.012 0 0 9.012 483.508 247.675 Tm -0.0002 Tc 0.0002 Tw ( = 2.03 *)Tj -46.6061 -1.4814 TD -0.0001 Tc -0.0014 Tw [(Homoptera - Aphidae)-10205.9(90.61)-1808.7(\(23.41\))-7983.1(77.44)-1805.6(\()-0.5(32.69\))-4806.4(t)]TJ 6.008 0 0 6.008 477.47 232.821 Tm 0.0001 Tc (42)Tj 9.012 0 0 9.012 483.508 234.324 Tm -0.0009 Tc 0.0009 Tw [( = 2.02 n.)-5.7(s.)]TJ -46.6061 -1.4814 TD 0.0002 Tc [(Delphacidae)-14423.3(4)0.1(.29)-2305.4(\()-0.2(1.39\))-8486(7)0.1(.48)-1802.1(\()-0.2(19.41\))-4806.1(t)]TJ 6.008 0 0 6.008 477.47 219.47 Tm 0.0001 Tc (42)Tj 9.012 0 0 9.012 483.508 220.972 Tm -0.0009 Tc [( = 2.02 n.)-5.7(s.)]TJ -46.6061 -1.4814 TD -0.0003 Tc [(Cicadellidae)-13926.9(10.62)-2309(\(1.79\))-7983.3(15.24)-2309(\()-0.7(3.81\))-4806.6(t)]TJ 6.008 0 0 6.008 477.47 206.119 Tm 0.0001 Tc (42)Tj 9.012 0 0 9.012 483.508 207.621 Tm -0.0003 Tc 0.0003 Tw ( = 2.02 n.s.)Tj -46.6061 -1.4814 TD -0.0001 Tc 0.0017 Tw [(Other Auchenorrhyncha)-9734.1(2.37)-2305.7(\(0.68\))-8486.3(7.95)-2305.7(\(1.34\))-4806.4(t)]TJ 6.008 0 0 6.008 477.47 192.768 Tm 0.0001 Tc (36)Tj 9.012 0 0 9.012 483.508 194.27 Tm -0.0001 Tc 0.0001 Tw ( = 2.03 ***)Tj -46.6061 -1.4814 TD 0.0017 Tw [(Small diptera)-13561.8(57.82)-2308.8(\(7.82\))-7486.2(110.85)-1802.4(\(13.78\))-4806.4(t)]TJ 6.008 0 0 6.008 477.47 179.417 Tm 0.0001 Tc (42)Tj 9.012 0 0 9.012 483.508 180.919 Tm -0.0002 Tc 0.0002 Tw ( = 2.02 *)Tj -46.6061 -1.4814 TD 0 Tc 0.0016 Tw [(Hymenopteran larvae)-10787.6(3.52)-2305.6(\(0.75\))-8486.2(3.24)-2305.6(\(1.21\))-4806.3(t)]TJ 6.008 0 0 6.008 477.47 166.065 Tm 0.0001 Tc (42)Tj 9.012 0 0 9.012 483.508 167.568 Tm -0.0009 Tc 0.0009 Tw [( = 2.02 n.)-5.7(s.)]TJ -46.6061 -1.4814 TD 0.0001 Tc [(Formicidae)-14866.9(0.55)-2305.5(\(0.28\))-8486.1(5.82)-2305.5(\(2.23\))-4806.2(t)]TJ 6.008 0 0 6.008 477.47 152.714 Tm (24)Tj 9.012 0 0 9.012 483.508 154.217 Tm -0.0002 Tc 0.0002 Tw ( = 2.06 *)Tj -46.6061 -1.4814 TD -0.0001 Tc 0.0017 Tw [(Lepidopteran larvae)-11401.1(2.49)-2305.7(\(1.20\))-8486.3(3.38)-2305.7(\(1.21\))-4806.4(t)]TJ 6.008 0 0 6.008 477.47 139.363 Tm 0.0001 Tc (42)Tj 9.012 0 0 9.012 483.508 140.865 Tm -0.0009 Tc 0.0009 Tw [( = 2.02 n.)-5.7(s.)]TJ -46.6061 -1.4751 TD 0 Tc [(Dermaptera)-14703.5(0.65)-2305.6(\(0.26\))-8486.2(0.67)-2305.6(\(0.48\))-4806.3(t)]TJ 6.008 0 0 6.008 477.47 126.069 Tm 0.0001 Tc (37)Tj 9.012 0 0 9.012 483.508 127.571 Tm -0.0009 Tc [( = 2.03 n.)-5.7(s.)]TJ -46.6061 -1.4814 TD 0 Tc [(Linyphiidae)-14590.2(2.05)-2305.6(\(0.88\))-8486.2(3.90)-2305.6(\(1.11\))-4806.3(t)]TJ 6.008 0 0 6.008 477.47 112.717 Tm 0.0001 Tc (42)Tj 9.012 0 0 9.012 483.508 114.22 Tm -0.0009 Tc [( = 2.02 n.)-5.7(s.)]TJ -46.6061 -1.4814 TD -0.0004 Tc 0.002 Tw [(Other araneae)-13845.2(4)-0.5(.37)-2306(\()-0.8(0.87\))-7983.4(1)-3.6(6.39)-2309.1(\(3.01\))-4806.7(t)]TJ 6.008 0 0 6.008 477.47 99.366 Tm 0.0001 Tc (35)Tj 9.012 0 0 9.012 483.508 100.869 Tm -0.0001 Tc 0.0001 Tw ( = 2.03 ***)Tj -46.6061 -1.4814 TD 0.0001 Tc [(Opiliones)-15532.1(0)0(.38)-2305.5(\()-0.3(0.23\))-8486.1(0)0(.45)-2305.5(\()-0.3(0.16\))-4806.2(t)]TJ 6.008 0 0 6.008 477.47 86.015 Tm (42)Tj 9.012 0 0 9.012 483.508 87.517 Tm -0.0009 Tc 0.0009 Tw [( = 2.02 n.)-5.7(s.)]TJ /F9 1 Tf 11 0 0 11 59.502 403.297 Tm 0 0 0 1 k /GS2 gs 0.0014 Tc 0 Tw ( )Tj /F14 1 Tf 10 0 0 10 65.143 403.297 Tm -0.0046 Tc 0.0136 Tw [(T)89(able 2. Mean chickfood invertebrate catch, per 100 samples, in beetle banks \(bb\) and field mar)29.4(gins \(fm\) sampled in)]TJ -0.5641 -1.1339 TD -0.0041 Tc 0.0122 Tw (June/July 1999, n = 22. Results of )Tj /F16 1 Tf 13.8954 0 TD 0 Tc 0 Tw (t)Tj /F14 1 Tf 0.275 0 TD -0.0044 Tc 0.015 Tw (-test on log)Tj /F15 1 Tf 5.8 0 0 5.8 246.107 389.549 Tm -0.0049 Tc (10)Tj /F14 1 Tf 10 0 0 10 251.918 391.958 Tm -0.0044 Tc 0.0125 Tw [( \(x + 1\) transformed data. \(*)]TJ /F16 1 Tf 11.503 0 TD 0 Tc 0 Tw (P)Tj /F14 1 Tf 0.5499 0 TD -0.0036 Tc 0.0133 Tw [( < 0.05, **)]TJ /F16 1 Tf 4.4362 0 TD 0 Tc 0 Tw (P)Tj /F14 1 Tf 0.5499 0 TD -0.0041 Tc 0.0116 Tw [( < 0.01, ***)]TJ /F16 1 Tf 4.9238 0 TD 0 Tc 0 Tw (P)Tj /F14 1 Tf 0.5499 0 TD -0.0057 Tc 0.015 Tw [( < 0.001\))]TJ ET /GS1 gs 59.502 380.761 m 530.053 380.761 l 59.502 366.305 m 530.053 366.305 l 59.502 82.557 m 530.053 82.557 l S BT /F2 1 Tf 9.01 0 0 9.01 217.363 732.201 Tm 0 g -0.0005 Tc 0.0022 Tw [(May mean)-2524(\()]TJ 6.998 0 0 6.998 281.426 732.201 Tm -0.0011 Tc (SE)Tj 9.01 0 0 9.01 289.59 732.201 Tm 6.727 Tc -6.7253 Tw [(\)A)6722.2(u)6726.8(g)6729.9( m)6727.9(e)6724.2(a)6727.4(n)4203.5(\()]TJ 6.998 0 0 6.998 416.271 732.201 Tm -0.0011 Tc (SE)Tj 9.01 0 0 9.01 424.434 732.201 Tm 0 Tc 0 Tw (\))Tj -39.9153 -1.8153 TD -0.0001 Tc [(Carabidae)-13930.9(1)-3.5(8.75)-3253.5(\(7.01\))-7044.8(29.04)-3256.6(\(6.28\))-4811.1(t)]TJ 6.007 0 0 6.007 478.945 714.342 Tm 0.0002 Tc (16)Tj 9.01 0 0 9.01 484.982 715.845 Tm -0.0007 Tc 0.0008 Tw [( = 2.12)-4.1( n.s.)]TJ -46.6355 -1.4818 TD 0.0002 Tc [(Staphylinidae)-12483.4(81.46)-2752.9(\()-0.3(25.39\))-6544.3(157.69)-2756.1(\(37.14\))-4810.8(t)]TJ 6.007 0 0 6.007 478.945 701.02 Tm (16)Tj 9.01 0 0 9.01 484.982 702.494 Tm -0.0007 Tc [( = 2.12)-4.1( n.s.)]TJ -46.6355 -1.4787 TD 0.0001 Tc [(Chrysomelidae)-12430(9.58)-3253.3(\(3.28\))-7044.6(25.58)-3256.4(\(8.34\))-4810.9(t)]TJ 6.007 0 0 6.007 478.945 687.668 Tm 0.0002 Tc (16)Tj 9.01 0 0 9.01 484.982 689.171 Tm -0.0007 Tc [( = 2.12)-4.1( n.s.)]TJ -46.6355 -1.4787 TD -0.0004 Tc 0.0021 Tw [(Cu)-3.8(rcurlion)-3.8(i)0.7(dae)-12487.1(3)-0.6(.75)-3253.8(\()-4(1.50\))-7045.1(14.62)-3008.4( \(8.47\))-4811.4(t)]TJ 6.007 0 0 6.007 478.945 674.346 Tm 0.0002 Tc (16)Tj 9.01 0 0 9.01 484.982 675.848 Tm -0.0007 Tc 0.0008 Tw [( = 2.12)-4.1( n.s.)]TJ -46.6355 -1.4818 TD 0 Tc 0.0017 Tw [(Elateridae)-14434.2(2.11)-3004.8( \(0.69\))-8794(0)-10483.2(t)]TJ 6.007 0 0 6.007 478.945 660.994 Tm 0.0002 Tc (16)Tj 9.01 0 0 9.01 484.982 662.497 Tm -0.0002 Tw ( = 2.12 **)Tj -46.6355 -1.4818 TD 0.0001 Tc [(Coccinellidae)-12486.6(2)-3.3(6.67)-2753(\(17.80\))-7541.7(3.08)-3256.4(\(1.19\))-4810.9(t)]TJ 6.007 0 0 6.007 478.945 647.643 Tm 0 Tc 0 Tw (9)Tj 9.01 0 0 9.01 481.949 649.146 Tm -0.0006 Tc 0.003 Tw [( = 2.26 n.s.)]TJ -46.2989 -1.4787 TD 0.0005 Tc [(Nitidulidae)-13983.8(0.42)-3252.9(\(0.27\))-8793.5(0)-10482.7(t)]TJ 6.007 0 0 6.007 478.945 634.32 Tm 0.0002 Tc (16)Tj 9.01 0 0 9.01 484.982 635.823 Tm -0.0007 Tc 0.0008 Tw [( = 2.12)-4.1( n.s.)]TJ -46.6355 -1.4818 TD 0 Tc [(Heteroptera)-13263.8(3)-3.4(5.83)-2753.1(\(17.45\))-7044.7(44.62)-2756.3(\(21.93\))-4811(t)]TJ 6.007 0 0 6.007 478.945 620.969 Tm 0.0002 Tc (16)Tj 9.01 0 0 9.01 484.982 622.472 Tm -0.0007 Tc [( = 2.12)-4.1( n.s.)]TJ -46.6355 -1.4818 TD 0 Tc -0.0015 Tw [(Homoptera - Aphidae)-9265(1)-3.4(5.94)-3253.4(\(5.78\))-7044.7(15.38)-3256.5(\(8.85\))-4811(t)]TJ 6.007 0 0 6.007 478.945 607.618 Tm 0.0002 Tc (16)Tj 9.01 0 0 9.01 484.982 609.12 Tm -0.0007 Tc 0.0008 Tw [( = 2.12)-4.1( n.s.)]TJ -46.6355 -1.4818 TD 0.0001 Tc [(Delphacidae)-12983.7(8)-3.3(4.08)-2753(\(41.70\))-6544.4(119.04)-2756.2(\(22.02\))-4810.9(t)]TJ 6.007 0 0 6.007 478.945 594.267 Tm 0 Tc 0 Tw (9)Tj 9.01 0 0 9.01 481.949 595.769 Tm -0.0006 Tc 0.003 Tw [( = 2.26 n.s.)]TJ -46.2989 -1.4818 TD -0.0001 Tc [(Cicadellidae)-12987(10.20)-3253.5(\()-3.7(3.12\))-7044.8(58.85)-2756.4(\(13.35\))-4811.1(t)]TJ 6.007 0 0 6.007 478.945 580.944 Tm 0.0002 Tc (16)Tj 9.01 0 0 9.01 484.982 582.418 Tm -0.0007 Tc 0.0008 Tw [( = 2.12)-4.1( n.s.)]TJ -46.6355 -1.4787 TD 0 Tc 0.0017 Tw [(Other Auchenorrhyncha)-10045.3(0)-13214.1(1)-0.2(.15)-3256.5(\()-0.5(0.82\))-4811(t)]TJ 6.007 0 0 6.007 478.945 567.593 Tm 0.0002 Tc (16)Tj 9.01 0 0 9.01 484.982 569.095 Tm 0 Tc 0.0001 Tw ( = 2.12 ***)Tj -46.6355 -1.4818 TD 0.0001 Tc 0.0016 Tw [(Small diptera)-12621.9(8)-3.3(4.79)-2753(\(12.07\))-6544.4(670.19)-2252.8(\(150.44\))-4810.9(t)]TJ 6.007 0 0 6.007 478.945 554.242 Tm 0.0002 Tc (16)Tj 9.01 0 0 9.01 484.982 555.744 Tm 0 Tc 0.0001 Tw ( = 2.12 ***)Tj -46.6355 -1.4818 TD 0.0003 Tc 0.0014 Tw [(Hymenopteran larvae)-9846.8(9.38)-3253.1(\(7.84\))-7541.5(2.50)-3256.2(\(0.81\))-4810.7(t)]TJ 6.007 0 0 6.007 478.945 540.891 Tm (10)Tj 9.01 0 0 9.01 484.982 542.393 Tm -0.0007 Tc 0.0008 Tw [( = 2.23)-4.1( n.s.)]TJ -46.6355 -1.4818 TD 0.0003 Tc [(Formicidae)-13427.1(34.38)-2752.8(\()-0.2(13.63\))-7044.4(40.19)-2756(\(10.04\))-4810.7(t)]TJ 6.007 0 0 6.007 478.945 527.539 Tm (16)Tj 9.01 0 0 9.01 484.982 529.042 Tm -0.0007 Tc [( = 2.12)-4.1( n.s.)]TJ -46.6355 -1.4787 TD 0.0002 Tc 0.0015 Tw [(Lepidopteran larvae)-10460.4(0.49)-3253.2(\(0.33\))-7541.6(3.46)-3256.3(\(1.18\))-4810.8(t)]TJ 6.007 0 0 6.007 478.945 514.216 Tm (16)Tj 9.01 0 0 9.01 484.982 515.719 Tm -0.0002 Tw ( = 2.12 *)Tj -46.6355 -1.4818 TD 0.0005 Tc [(Dermapte)4(ra)-13263.3(10.01)-3252.9(\()-3.1(7.01\))-7541.3(0)0.3(.19)-3256(\()0(0.19\))-4810.5(t)]TJ 6.007 0 0 6.007 478.945 500.865 Tm 0 Tc 0 Tw (8)Tj 9.01 0 0 9.01 481.949 502.368 Tm -0.0003 Tc 0.0027 Tw [( = 2.31 *)]TJ -46.2989 -1.4787 TD 0.0003 Tc [(Linyphiidae)-12650(154.38)-2752.8(\(26.09\))-6544.2(251.73)-2756(\(48.03\))-4810.7(t)]TJ 6.007 0 0 6.007 478.945 487.542 Tm (16)Tj 9.01 0 0 9.01 484.982 489.045 Tm -0.0007 Tc 0.0008 Tw [( = 2.12)-4.1( n.s.)]TJ -46.6355 -1.4818 TD -0.0002 Tc 0.0019 Tw [(Other araneae)-12405.1(3)-3.6(1.04)-3253.6(\(8.42\))-7044.9(42.88)-3256.7(\(5.81\))-4811.2(t)]TJ 6.007 0 0 6.007 478.945 474.191 Tm 0 Tc 0 Tw (9)Tj 9.01 0 0 9.01 481.949 475.694 Tm -0.0006 Tc 0.003 Tw [( = 2.26 n.s.)]TJ -46.2989 -1.4787 TD 0 Tc [(Opiliones)-14596.2(4)-0.2(.98)-3253.4(\()-3.6(3.29\))-7044.7(10.58)-3256.5(\(2.32\))-4811(t)]TJ 6.007 0 0 6.007 478.945 460.868 Tm 0.0002 Tc (16)Tj 9.01 0 0 9.01 484.982 462.371 Tm -0.0002 Tw ( = 2.12 *)Tj ET endstream endobj 24 0 obj << /ProcSet [/PDF /Text ] /Font << /F2 25 0 R /F4 4 0 R /F5 5 0 R /F6 6 0 R /F9 9 0 R /F14 26 0 R /F15 27 0 R /F16 28 0 R >> /ExtGState << /GS1 29 0 R /GS2 13 0 R >> >> endobj 31 0 obj << /Length 17266 >> stream BT /F4 1 Tf 9 0 0 9 522.205 788.95 Tm 0 0 0 1 k /GS2 gs -0.0019 Tc (115)Tj /F5 1 Tf 8 0 0 8 230.038 788.95 Tm -0.0057 Tc 0.0954 Tw [(Resour)29.7(ce pr)29.7(ovision for farmland gamebir)33.3(d)8.3(s)]TJ /F9 1 Tf 11 0 0 11 65.147 762.985 Tm -0.0062 Tc 0.0727 Tw [(habitats \(T)60.8(able 2\).)]TJ 0.822 -1.09 TD 0.0114 Tw (The diversity \(H)Tj /F17 1 Tf 6.5429 0 TD 0 Tc 0 Tw (\242)Tj /F9 1 Tf 0.2448 0 TD -0.0058 Tc 0.0114 Tw (\) of chickfood invertebrates was)Tj -7.6097 -1.09 TD -0.006 Tc 0.0154 Tw [(significantly higher in field mar)12(gins \(F)]TJ /F10 1 Tf 6.4 0 0 6.4 234.092 735.886 Tm -0.0017 Tc 0.0156 Tw (1, 34)Tj 2.0153 -0.6644 TD 0 Tc 0 Tw ( )Tj /F12 1 Tf 11 0 0 11 248.69 739.004 Tm -0.0109 Tc (= )Tj /F9 1 Tf 0.8246 0 TD -0.0052 Tc (5.20, )Tj /F11 1 Tf 2.2394 0 TD 0 Tc (P)Tj /F9 1 Tf -19.7497 -1.0926 TD -0.0058 Tc 0.0636 Tw (= 0.03\), with a mean index of 1.51 compared with)Tj 0 -1.09 TD -0.0055 Tc 0.075 Tw (1.40 in beetle banks \()Tj /F13 1 Tf 8.8 0 0 8.8 161.553 714.994 Tm -0.0082 Tc (SED)Tj /F9 1 Tf 11 0 0 11 177.994 714.994 Tm -0.0059 Tc 0.0754 Tw [( = 0.08\). However)32.8(, it did)]TJ -10.2588 -1.09 TD -0.0061 Tc 0.0372 Tw [(not dif)11.9(fer significantly between farms \(F)]TJ /F10 1 Tf 6.4 0 0 6.4 243.049 699.886 Tm -0.002 Tc 0.0391 Tw (4, 34)Tj 2.0374 -0.6644 TD 0 Tc 0 Tw ( )Tj /F12 1 Tf 11 0 0 11 257.959 703.004 Tm -0.0098 Tc (= )Tj /F9 1 Tf 0.8478 0 TD -0.0059 Tc (0.56,)Tj /F11 1 Tf -18.3762 -1.0926 TD 0 Tc (P)Tj /F9 1 Tf 0.6056 0 TD -0.0056 Tc 0.0386 Tw [( = 0.69\), nor was there any significant interaction)]TJ -0.6056 -1.0901 TD -0.0061 Tc -0.0045 Tw (between these habitat and farm factors \(F)Tj /F10 1 Tf 6.4 0 0 6.4 244.155 675.905 Tm -0.002 Tc -0.0039 Tw (4, 34 )Tj /F12 1 Tf 11 0 0 11 258.498 678.994 Tm -0.0114 Tc (= )Tj /F9 1 Tf 0.7989 0 TD -0.0059 Tc (1.39,)Tj /F11 1 Tf -18.3762 -1.09 TD 0 Tc 0 Tw (P)Tj /F9 1 Tf 0.6056 0 TD -0.0059 Tc 0.0091 Tw [( = 0.29\).)]TJ /F11 1 Tf 1.6441 -2.1827 TD 0.0499 Tw [(Summer plant pr)30.2(ovision in beetle banks)]TJ /F9 1 Tf -1.4276 -1.09 TD -0.0061 Tc -0.0442 Tw (1998 sampling revealed that there was more cover)Tj -0.822 -1.0926 TD 0.0499 Tw [(in the field mar)11.8(gin bases compared to beetle banks)]TJ 0 -1.0901 TD 0.0025 Tc (\(t)Tj /F10 1 Tf 6.4 0 0 6.4 72.092 603.905 Tm 0.0133 Tc (16)Tj /F9 1 Tf 11 0 0 11 78.753 606.994 Tm 0.0125 Tc 0.1228 Tw [( = 1.75, )]TJ /F11 1 Tf 3.7984 0 TD 0 Tc 0 Tw (P)Tj /F9 1 Tf 0.6236 0 TD 0.0124 Tc 0.1227 Tw [( = 0.003\) \(Fig. 1\); however)35.6(, cover)]TJ -5.659 -1.09 TD -0.0053 Tc -0.0046 Tw (provision was high in both kinds of habitat, with no)Tj 0 -1.0926 TD -0.0065 Tc -0.0282 Tw [(less than 67% cover)32.2(, and a maximum of 97%. )14.2(There)]TJ 0 -1.09 TD -0.0047 Tc 0.1227 Tw [(was no significant dif)13.3(ference in the percentage of)]TJ T* -0.0094 Tc -0.0614 Tw (tussocky-structured grass present in beetle banks and)Tj 0 -1.0926 TD -0.0069 Tc 0.0434 Tw [(field mar)11.1(gin \(t)]TJ /F10 1 Tf 6.4 0 0 6.4 128.444 543.895 Tm 0 Tc (12)Tj /F9 1 Tf 11 0 0 11 134.851 546.985 Tm -0.0058 Tc 0.0454 Tw [( = 1.78, )]TJ /F11 1 Tf 3.4042 0 TD 0 Tc 0 Tw (P)Tj /F9 1 Tf 0.6056 0 TD -0.0058 Tc 0.0455 Tw [( = 0.14\). Other grass and)]TJ -10.3465 -1.09 TD -0.0057 Tc 0.0029 Tw [(herbaceous plant cover)35.5(, fed on by adult game birds,)]TJ T* -0.0058 Tc 0.03 Tw [(was not significantly dif)12.2(ferent overall \(t)]TJ /F10 1 Tf 6.4 0 0 6.4 239.052 519.886 Tm 0 Tc (16)Tj /F9 1 Tf 11 0 0 11 245.459 523.004 Tm -0.0056 Tc 0.0303 Tw [( = 1.75, )]TJ /F11 1 Tf 3.3578 0 TD 0 Tc 0 Tw (P)Tj /F9 1 Tf -19.7497 -1.0926 TD -0.0061 Tc -0.0137 Tw (= 0.10, grasses; t)Tj /F10 1 Tf 6.4 0 0 6.4 137.912 507.895 Tm 0 Tc (10)Tj /F9 1 Tf 11 0 0 11 144.29 510.985 Tm -0.0059 Tc -0.0136 Tw [( = 1.75, )]TJ /F11 1 Tf 3.2289 0 TD 0 Tc 0 Tw (P)Tj /F9 1 Tf 0.603 0 TD -0.0055 Tc -0.0145 Tw [( = 0.07, dicots\) \(Fig. 1\).)]TJ -11.0268 -1.09 TD 0.0181 Tc 0.1227 Tw [(Field mar)7.8(gin bases contained small amounts of)]TJ T* -0.0057 Tc -0.0033 Tw (woody plants, the presence of which was negligible)Tj T* 0.0068 Tc 0.1228 Tw [(in beetle banks \(Fig. 1\). )9.5(There was considerable)]TJ 0 -1.0926 TD -0.0056 Tc 0.0181 Tw (variation in the abundance of these plant categories)Tj 0 -1.0901 TD -0.0058 Tc 0.0863 Tw (between individual fields.)Tj 0.822 -1.09 TD 0.0327 Tc 0.1229 Tw [(In 1999, there was no significant dif)14.7(ference)]TJ -0.822 -1.0926 TD -0.006 Tc 0.0205 Tw [(between overall vegetational cover in field mar)12(gins)]TJ 0 -1.09 TD -0.0058 Tc -0.0137 Tw (and beetle banks \(t)Tj /F10 1 Tf 6.4 0 0 6.4 146.303 411.914 Tm 0 Tc (37)Tj /F9 1 Tf 11 0 0 11 152.709 415.004 Tm -0.0059 Tc -0.0136 Tw [( = 1.69, )]TJ /F11 1 Tf 3.2263 0 TD 0 Tc 0 Tw (P)Tj /F9 1 Tf 0.6056 0 TD -0.0056 Tc -0.0137 Tw [( = 0.10\), and levels of)]TJ -11.7921 -1.09 TD -0.0058 Tc -0.0085 Tw (both tussocky grass and other grass species were no)Tj 0 -1.0926 TD -0.0063 Tc 0.0821 Tw [(dif)11.7(ferent either \(t)]TJ /F10 1 Tf 6.4 0 0 6.4 140.69 387.905 Tm 0 Tc (42)Tj /F9 1 Tf 11 0 0 11 147.096 390.994 Tm -0.0059 Tc 0.0834 Tw [( = 1.68, )]TJ /F11 1 Tf 3.5175 0 TD 0 Tc 0 Tw (P)Tj /F9 1 Tf 0.6056 0 TD -0.0061 Tc 0.0827 Tw [( = 0.1)30(1, tussock; t)]TJ /F10 1 Tf 6.4 0 0 6.4 272.898 387.905 Tm 0 Tc (34)Tj /F9 1 Tf 11 0 0 11 279.304 390.994 Tm 0.072 Tc 0 Tw ( =)Tj -19.4689 -1.09 TD -0.0053 Tc (1.69, )Tj /F11 1 Tf 2.3373 0 TD 0 Tc (P)Tj /F9 1 Tf 0.603 0 TD -0.0063 Tc 0.117 Tw [( = 0.69, grass\). )47.9(As expected, field mar)11.7(gins)]TJ -2.9403 -1.09 TD 0.008 Tc 0.1228 Tw (contained significantly more dicotyledonous and)Tj 0 -1.0926 TD -0.0102 Tc -0.0609 Tw (woody plants than beetle banks \(t)Tj /F10 1 Tf 6.4 0 0 6.4 205.689 351.905 Tm -0.0088 Tc (26)Tj /F9 1 Tf 11 0 0 11 212.038 354.995 Tm -0.0102 Tc -0.0606 Tw [( = 1.71, )]TJ /F11 1 Tf 3.046 0 TD 0 Tc 0 Tw (P)Tj /F9 1 Tf 0.6004 0 TD -0.0108 Tc -0.0614 Tw [( < 0.001,)]TJ -17.0001 -1.09 TD -0.0064 Tc 0.0273 Tw (dicots; t)Tj /F10 1 Tf 6.4 0 0 6.4 100.041 339.914 Tm 0 Tc (42)Tj /F9 1 Tf 11 0 0 11 106.448 343.004 Tm -0.0056 Tc 0.0303 Tw [( = 1.68, )]TJ /F11 1 Tf 3.3603 0 TD 0 Tc 0 Tw (P)Tj /F9 1 Tf 0.603 0 TD -0.0053 Tc 0.0303 Tw [( < 0.001, woody plants; Fig. 2\).)]TJ -7.718 -1.09 TD -0.0059 Tc 0.0864 Tw (In the majority of vegetation categories, there was)Tj 0 -1.0926 TD -0.0062 Tc 0.0834 Tw (considerable variation between farms.)Tj /F11 1 Tf 2.5538 -2.1801 TD -0.0058 Tc 0.0491 Tw [(W)58.8(inter plant pr)30.3(ovision in beetle banks)]TJ /F9 1 Tf -1.7317 -1.0926 TD 0.0372 Tc 0.1227 Tw [(There was no significant dif)14(ference in total)]TJ -0.822 -1.09 TD 0.0089 Tc (vegetation cover between beetle banks and field)Tj T* -0.0064 Tc 0.1 Tw [(m)6.6(a)6.6(r)11.6(gins \(t)]TJ /F10 1 Tf 6.4 0 0 6.4 110.246 255.895 Tm 0 Tc (16)Tj /F9 1 Tf 11 0 0 11 116.652 259.013 Tm -0.0059 Tc 0.1015 Tw [( = 1.75, )]TJ /F11 1 Tf 3.5717 0 TD 0 Tc 0 Tw (P)Tj /F9 1 Tf 0.6056 0 TD -0.0054 Tc 0.1015 Tw [( = 0.33\) \(Fig. 3\). Cover had)]TJ -8.8596 -1.0926 TD 0.001 Tc 0.1228 Tw (been maintained with a very similar range to the)Tj 0 -1.09 TD -0.0068 Tc 0.0871 Tw [(summer)31.9(, at between 62% \226 97 %. However)34.4(, when)]TJ T* 0.0008 Tc 0.1227 Tw (mean percentage of tussocky live and dead grass)Tj 0 -1.0926 TD -0.0062 Tc 0.1053 Tw (cover was examined, there was significantly more)Tj 0 -1.0901 TD -0.0059 Tc 0.0789 Tw [(present in beetle banks than in field mar)12.1(gins \(t)]TJ /F10 1 Tf 6.4 0 0 6.4 272.955 195.914 Tm 0 Tc (16)Tj /F9 1 Tf 11 0 0 11 279.361 199.004 Tm 0.0669 Tc 0 Tw ( =)Tj -19.474 -1.09 TD 0.0073 Tc (1.75, )Tj /F11 1 Tf 2.4172 0 TD 0 Tc (P)Tj /F9 1 Tf 0.6185 0 TD 0.0068 Tc 0.1228 Tw [( = 0.01\). Mean percentage of other grass)]TJ -3.0356 -1.0926 TD -0.0057 Tc 0.0344 Tw [(growing in the habitat was not dif)12.3(ferent \(t)]TJ /F10 1 Tf 6.4 0 0 6.4 248.605 171.905 Tm 0 Tc (16)Tj /F9 1 Tf 11 0 0 11 255.011 174.995 Tm -0.006 Tc 0.0363 Tw [( = 1.75,)]TJ /F11 1 Tf -17.2604 -1.09 TD 0 Tc 0 Tw (P)Tj /F9 1 Tf 0.6649 0 TD 0.0557 Tc 0.1253 Tw [( = 0.12\), but there was significantly more)]TJ -0.6649 -1.09 TD -0.0061 Tc -0.0075 Tw [(dicotyledonous plant material in field mar)11.9(gins \(t)]TJ /F10 1 Tf 6.4 0 0 6.4 273.947 147.895 Tm 0 Tc (16)Tj /F9 1 Tf 11 0 0 11 280.353 151.013 Tm -0.0233 Tc 0 Tw ( =)Tj -19.5642 -1.0926 TD -0.0054 Tc (1.75, )Tj /F11 1 Tf 2.2548 0 TD 0 Tc (P)Tj /F9 1 Tf 0.6056 0 TD -0.0059 Tc 0.0328 Tw [( = 0.01\), as well as more woody plants \(t)]TJ /F10 1 Tf 6.4 0 0 6.4 276.64 135.905 Tm 0 Tc 0 Tw (8)Tj /F9 1 Tf 11 0 0 11 279.843 138.995 Tm 0.0231 Tc ( =)Tj -19.5178 -1.09 TD -0.0052 Tc (1.86, )Tj /F11 1 Tf 2.2419 0 TD 0 Tc (P)Tj /F9 1 Tf 0.603 0 TD -0.0054 Tc 0.0182 Tw [( = 0.02\) \(Fig. 3\).)]TJ -2.0229 -1.09 TD -0.0061 Tc -0.0499 Tw [(Plant litter)32.6(, dead grass and tussock were combined)]TJ -0.822 -1.0926 TD 0.1045 Tw (into a single category that was compared between)Tj 0 -1.0901 TD -0.0033 Tc 0.1226 Tw [(habitats. )14.6(These vegetation types are important for)]TJ T* -0.0097 Tc -0.0613 Tw (providing camouflaged nesting material in the spring)Tj 22.422 62.1792 TD -0.0057 Tc -0.01 Tw (\(Rands, 1988\). Beetle banks contained significantly)Tj 0 -1.09 TD -0.0066 Tc 0.0943 Tw (more of this material \(t)Tj /F10 1 Tf 6.4 0 0 6.4 415.707 747.905 Tm 0 Tc (16)Tj /F9 1 Tf 11 0 0 11 422.085 750.994 Tm -0.0059 Tc 0.0955 Tw [( = 1.75, )]TJ /F11 1 Tf 3.5562 0 TD 0 Tc 0 Tw (P)Tj /F9 1 Tf 0.603 0 TD -0.0058 Tc 0.0954 Tw [( = 0.01\) with a)]TJ -14.1861 -1.09 TD -0.0064 Tc 0.0327 Tw (mean of 61% compared to 27%.)Tj ET q 1 i 311.789 266.157 223.909 -115.398 re W n 1 g /GS1 gs 311.789 266.157 287.093 -138.387 re f Q 0.754 g 0 G 1 J 0 j 0.515 w 10 M []0 d /GS1 gs 1 i 372.224 195.546 38.381 -39.43 re B* 467.95 164.024 37.871 -7.909 re B* 0 g 372.224 238.632 38.381 -43.087 re B* 467.95 214.963 37.871 -50.939 re B* 1 g 467.95 230.752 37.871 -15.789 re B* 0.5 g 372.224 240.73 38.381 -2.098 re B* 467.95 245.974 37.871 -15.222 re B* 1 j 0.02 w 343.821 260.176 m 343.849 156.116 l 341.185 156.116 m 343.821 156.087 l 341.185 177.149 m 343.821 177.092 l 341.185 197.643 m 343.821 197.587 l 341.185 218.648 m 343.821 218.62 l 341.185 239.143 m 343.821 239.114 l 341.185 260.176 m 343.821 260.147 l 343.821 156.116 m 534.734 156.087 l 343.821 153.508 m 343.849 156.116 l 439.547 153.508 m 439.575 156.116 l 534.734 153.508 m 534.762 156.116 l S BT /F2 1 Tf 9.815 0 0 9.811 333.304 151.722 Tm 0 g 0 Tc 0 Tw (0)Tj -0.4303 2.1408 TD 0.0025 Tc (20)Tj 0 2.0888 TD (40)Tj 0 2.1408 TD (60)Tj 0 2.0888 TD (80)Tj -0.4275 2.1437 TD 0.0011 Tc (100)Tj ET q 311.789 222.418 8.787 -27.808 re W* n BT 0 8.404 -8.401 0 318.621 194.582 Tm 0.0007 Tc -0.0011 Tw (% cover)Tj ET Q BT /F18 1 Tf 9.7 0 0 9.7 366.441 140.015 Tm 0 0 0 1 k /GS2 gs -0.0059 Tc 0.0777 Tw [(Beetle banks)-4906.6(Field )-14.4(mar)8.7(gins)]TJ /F16 1 Tf 10 0 0 10 317.459 123.602 Tm -0.0044 Tc 0.0763 Tw (Fig. 3. Mean percentage cover of plant categories in)Tj 0 -1.0998 TD 0.036 Tc 0.125 Tw [(beetle banks and field mar)10.5(gins, 1998-99 winter)]TJ T* -0.004 Tc -0.0331 Tw (assessment; n = 18. Light grey shading: tussock grass;)Tj T* 0.0007 Tc 0.125 Tw [(Black: other grasses; )14.9(White: dicotyledonous plants;)]TJ T* -0.0036 Tc -0.0034 Tw (Dark grey: woody plants.)Tj ET 0.754 g 0 j 0.515 w /GS1 gs 371.884 651.328 38.409 -58.337 re B* 467.638 632.421 38.381 -39.43 re B* 0 g 371.884 668.705 38.409 -17.376 re B* 467.638 661.873 38.381 -29.452 re B* 1 g 371.884 678.683 38.409 -9.978 re B* 467.638 685.004 38.381 -23.131 re B* 0.5 g 371.884 679.221 38.409 -0.539 re B* 467.638 692.346 38.381 -7.342 re B* 1 j 0.02 w 343.509 702.352 m 343.537 592.991 l 340.873 592.991 m 343.509 592.963 l 340.873 604.018 m 343.509 603.99 l 340.873 615.073 m 343.509 615.045 l 340.873 625.59 m 343.509 625.561 l 340.873 636.645 m 343.509 636.588 l 340.873 647.672 m 343.509 647.615 l 340.873 658.698 m 343.509 658.67 l 340.873 669.725 m 343.509 669.725 l 340.873 680.27 m 343.509 680.242 l 340.873 691.297 m 343.509 691.268 l 340.873 702.352 m 343.509 702.295 l 343.509 592.991 m 534.932 592.963 l 343.509 590.355 m 343.537 592.991 l 439.207 590.355 m 439.263 592.991 l 534.932 590.355 m 534.961 592.991 l S BT /F2 1 Tf 9.815 0 0 9.815 332.992 588.569 Tm 0 g 0 Tc 0 Tw (0)Tj -0.4303 1.1234 TD -0.0004 Tc (10)Tj T* (20)Tj 0 1.0715 TD (30)Tj 0 1.1263 TD (40)Tj T* (50)Tj 0 1.1234 TD (60)Tj T* (70)Tj 0 1.0715 TD (80)Tj 0 1.1263 TD (90)Tj -0.4274 1.1235 TD 0.0011 Tc (100)Tj ET q 310.74 666.607 11.197 -32.457 re W* n BT 0 9.815 -9.815 0 319.641 634.094 Tm 0.0002 Tc 0.0011 Tw (% cover)Tj ET Q BT /F18 1 Tf 9.7 0 0 9.7 365.903 577.542 Tm 0 0 0 1 k /GS2 gs -0.0059 Tc 0.0777 Tw [(Beetle banks)-4614.4(Field )-14.4(mar)11.6(gins)]TJ /F16 1 Tf 10 0 0 10 317.459 560.79 Tm -0.0044 Tc 0.0763 Tw (Fig. 1. Mean percentage cover of plant categories in)Tj 0 -1.0998 TD 0.0545 Tc 0.1251 Tw [(beetle banks and field mar)14.8(gins, summer 1998)]TJ T* -0.004 Tc -0.0331 Tw (assessment; n = 18. Light grey shading: tussock grass;)Tj T* 0.0007 Tc 0.125 Tw [(Black: other grasses; )14.9(White: dicotyledonous plants;)]TJ T* -0.0036 Tc -0.0034 Tw (Dark grey: woody plants.)Tj ET 0 0 0 0 k /GS1 gs 322.8 694.8 20.7 -11.6 re f* 322.8 673.1 20.7 -11.6 re f* 322.8 652.85 20.7 -11.6 re f* 322.8 629.9 20.7 -11.6 re f* 322.8 607.5 20.7 -11.6 re f* BT 10 0 0 10 317.685 339.007 Tm 0 0 0 1 k /GS2 gs -0.0044 Tc 0.0763 Tw (Fig. 2. Mean percentage cover of plant categories in)Tj T* 0.0545 Tc 0.1251 Tw [(beetle banks and field mar)12(gins, summer 1999)]TJ T* -0.004 Tc -0.0331 Tw (assessment; n = 44. Light grey shading: tussock grass;)Tj T* 0.0007 Tc 0.125 Tw [(Black: other grasses; )12(White: dicotyledonous plants;)]TJ T* -0.0036 Tc -0.0034 Tw (Dark grey: woody plants.)Tj /F18 1 Tf 9.7 0 0 9.7 366.186 355.306 Tm -0.0059 Tc 0.0777 Tw [(Beetle banks)-4903.7(Field )-14.4(mar)8.7(gins)]TJ ET 0.754 g 0 j 0.515 w /GS1 gs 373.244 411.291 39.458 -39.969 re B* 471.607 397.599 39.43 -26.277 re B* 0 g 373.244 444.909 39.458 -33.619 re B* 471.607 427.051 39.43 -29.452 re B* 1 g 373.244 461.209 39.458 -16.299 re B* 471.607 457.524 39.43 -30.472 re B* 0.5 g 471.607 469.089 39.43 -11.565 re B* q 310.655 487.486 229.55 -120.728 re W n 1 j 0.02 w 343.821 482.78 m 343.821 371.322 l 341.185 371.322 m 343.821 371.294 l 341.185 382.349 m 343.821 382.32 l 341.185 393.404 m 343.821 393.347 l 341.185 404.969 m 343.821 404.913 l 341.185 415.996 m 343.821 415.968 l 341.185 427.051 m 343.821 427.023 l 341.185 438.078 m 343.821 438.05 l 341.185 449.133 m 343.821 449.105 l 341.185 460.698 m 343.821 460.67 l 341.185 471.725 m 343.821 471.697 l 341.185 482.78 m 343.821 482.724 l 343.821 371.322 m 540.488 371.294 l 343.821 368.686 m 343.821 371.322 l 442.155 368.686 m 442.183 371.322 l S BT /F2 1 Tf 9.814 0 0 9.815 333.304 366.872 Tm 0 g 0 Tc 0 Tw (0)Tj ET Q BT /F2 1 Tf 9.814 0 0 9.815 329.109 377.898 Tm 0 g -0.0003 Tc (10)Tj 0 1.1263 TD (20)Tj 0 1.1783 TD (30)Tj 0 1.1263 TD (40)Tj 0 1.1234 TD (50)Tj 0 1.1263 TD (60)Tj 0 1.1234 TD (70)Tj 0 1.1783 TD (80)Tj 0 1.1234 TD (90)Tj -0.4304 1.1263 TD (100)Tj ET q 311.052 445.987 11.197 -32.485 re W* n BT 0 9.814 -9.815 0 319.953 413.473 Tm 0.0003 Tc 0.001 Tw (% cover)Tj ET Q 0 0 0 1 K 0 J 0.51 w 343.736 702.409 m 343.736 590.157 l 340.646 592.594 m 534.791 592.594 l 344.048 482.695 m 344.048 368.062 l 340.646 371.039 m 536.236 371.039 l S 0 0 0 0 k 534.85 376.25 6.95 -8.2 re f* 534.791 371.35 m 534.791 368.091 l 344.048 260.204 m 344.048 152.459 l 340.788 155.889 m 534.536 155.889 l S 322.95 476.6 20.7 -11.6 re f* 322.95 454.05 20.7 -11.6 re f* 322.95 431.1 20.7 -11.6 re f* 322.95 407.85 20.7 -11.6 re f* 322.95 386.3 20.7 -11.6 re f* 442.098 370.755 m 442.098 367.892 l 439.547 156.144 m 439.547 153.31 l 534.791 156.144 m 534.791 153.31 l 534.791 592.85 m 534.791 589.987 l 439.292 592.85 m 439.292 589.987 l 340.391 702.153 m 343.792 702.153 l 340.391 680.043 m 343.792 680.043 l 340.391 658.387 m 343.792 658.387 l 340.391 636.645 m 343.792 636.645 l 340.391 615.045 m 343.792 615.045 l 340.703 482.497 m 344.104 482.497 l 340.703 460.698 m 344.104 460.698 l 340.703 437.993 m 344.104 437.993 l 340.703 415.854 m 344.104 415.854 l 340.703 393.149 m 344.104 393.149 l 340.703 259.949 m 344.104 259.949 l 340.703 238.944 m 344.104 238.944 l 340.703 218.563 m 344.104 218.563 l 340.703 197.558 m 344.104 197.558 l 340.703 176.894 m 344.104 176.894 l S endstream endobj 32 0 obj << /ProcSet [/PDF /Text ] /Font << /F2 25 0 R /F4 4 0 R /F5 5 0 R /F9 9 0 R /F10 10 0 R /F11 11 0 R /F12 12 0 R /F13 21 0 R /F16 28 0 R /F17 33 0 R /F18 34 0 R >> /ExtGState << /GS1 29 0 R /GS2 13 0 R >> >> endobj 36 0 obj << /Length 12300 >> stream BT /F4 1 Tf 9 0 0 9 59.502 788.95 Tm 0 0 0 1 k /GS2 gs -0.0019 Tc (116)Tj /F6 1 Tf 8 0 0 8 258.948 788.95 Tm -0.0084 Tc 0.0207 Tw [(S R )11.3(THOMAS )]TJ /F5 1 Tf 6.1618 0 TD -0.0051 Tc 0.0052 Tw [(E)5.3(T)12.6( AL.)]TJ /F11 1 Tf 11 0 0 11 86.006 762.985 Tm -0.0057 Tc 0.0613 Tw (Relationships between plant cover and)Tj 5.1178 -1.09 TD 0.0021 Tc (invertebrates)Tj /F9 1 Tf -6.7052 -1.09 TD -0.006 Tc 0.1187 Tw (As the ages of the beetle banks assessed in this)Tj -0.822 -1.0926 TD -0.0059 Tc -0.0324 Tw (study were known, it was possible to assess whether)Tj 0 -1.09 TD -0.0055 Tc -0.0129 Tw (abundance and diversity of chick food invertebrates)Tj T* -0.0058 Tc 0.103 Tw (and plant cover showed age-related changes. Data)Tj 0 -1.0926 TD 0.0077 Tc 0.1227 Tw (from the second sampling year was used, which)Tj 0 -1.0901 TD -0.0054 Tc (represented a greater sample size of beetle banks.)Tj T* -0.0058 Tc -0.021 Tw (The two newly sown banks were excluded, as being)Tj 0 -1.0926 TD 0.0087 Tc 0.1227 Tw (less than a year old, they contained only annual)Tj 0 -1.09 TD -0.0056 Tc 0.0681 Tw (species that had germinated through the ploughing)Tj T* 0.061 Tw (disturbance whilst creating the bank, prior to grass)Tj 0 -1.0926 TD 0.1227 Tw (establishment. Regression analysis was performed)Tj 0 -1.0901 TD -0.006 Tc 0.1136 Tw (between age in years and mean invertebrate catch)Tj T* -0.0056 Tc 0.054 Tw (in beetle banks, but no relationship was evident \(r)Tj /F10 1 Tf 6.4 0 0 6.4 280.208 598.661 Tm 0 Tc 0 Tw (2)Tj /F9 1 Tf 11 0 0 11 59.502 582.985 Tm 0.0199 Tc 0.1227 Tw (< 0.001, F)Tj /F10 1 Tf 6.4 0 0 6.4 109.817 579.895 Tm 0.0252 Tc 0.125 Tw (1, 18)Tj 2.2544 -0.6644 TD 0 Tc 0 Tw ( )Tj /F9 1 Tf 11 0 0 11 126.797 582.985 Tm 0.0205 Tc 0.1227 Tw (= 0.015, )Tj /F11 1 Tf 3.9969 0 TD 0 Tc 0 Tw (P)Tj /F9 1 Tf 0.6314 0 TD 0.0202 Tc 0.1226 Tw [( = 0.905\). However)38.2(,)0.5( a)]TJ -10.7459 -1.09 TD -0.001 Tc 0.1227 Tw (regression between age and invertebrate diversity)Tj T* -0.0112 Tc -0.0607 Tw [(\(Shannon-W)32.6(iener Index\) showed a highly significant)]TJ 0 -1.0926 TD -0.0057 Tc 0.0636 Tw (positive relationship \(r)Tj /F10 1 Tf 6.4 0 0 6.4 159.055 550.642 Tm 0 Tc 0 Tw (2)Tj /F9 1 Tf 11 0 0 11 162.258 546.985 Tm -0.0057 Tc 0.0652 Tw [( = 0.335, F)]TJ /F10 1 Tf 6.4 0 0 6.4 211.864 543.895 Tm -0.0012 Tc 0.0625 Tw (1, 18)Tj 2.0595 -0.6644 TD 0 Tc 0 Tw ( )Tj /F9 1 Tf 11 0 0 11 227.115 546.985 Tm -0.0057 Tc 0.0637 Tw (= 9.084, )Tj /F11 1 Tf 3.6438 0 TD 0 Tc 0 Tw (P)Tj /F9 1 Tf 0.6056 0 TD 0.054 Tc ( =)Tj -19.4869 -1.09 TD 0.0557 Tc 0.3008 Tw [(0.008\). )14.4(This undoubtedly results from the)]TJ T* 0.0444 Tc 0.1228 Tw (increasingly complex plant communities that)Tj 0 -1.0926 TD -0.006 Tc 0.0281 Tw (develops in the beetle banks \(Thomas )Tj /F11 1 Tf 15.2813 0 TD -0.0065 Tc 0.0272 Tw (et al.)Tj /F9 1 Tf 1.9971 0 TD -0.0054 Tc 0.0319 Tw (, 2000\).)Tj -17.2785 -1.09 TD -0.0057 Tc -0.0324 Tw (Plant diversity was also calculated, and had a highly)Tj T* 0.004 Tc 0.1228 Tw (significant positive relationship with invertebrate)Tj T* -0.0045 Tc 0.1231 Tw [(diversity)57.4(, both in beetle banks \(r)]TJ /F10 1 Tf 6.4 0 0 6.4 206.365 478.642 Tm 0 Tc 0 Tw (2)Tj /F9 1 Tf 11 0 0 11 209.568 475.013 Tm -0.0044 Tc 0.1227 Tw [( = 0.559, F)]TJ /F10 1 Tf 6.4 0 0 6.4 261.215 471.895 Tm 0.0009 Tc 0.125 Tw (1, 18)Tj 2.1304 -0.6644 TD 0 Tc 0 Tw ( )Tj /F9 1 Tf 11 0 0 11 277.316 475.013 Tm (=)Tj -19.8013 -1.0926 TD -0.005 Tc (22.790, )Tj /F11 1 Tf 3.2238 0 TD 0 Tc (P)Tj /F9 1 Tf 0.603 0 TD -0.006 Tc 0.0091 Tw [( < 0.001\) and in field mar)12(gins \(r)]TJ /F10 1 Tf 6.4 0 0 6.4 241.316 466.651 Tm 0 Tc 0 Tw (2)Tj /F9 1 Tf 11 0 0 11 244.491 462.994 Tm -0.0059 Tc 0.0091 Tw [( = 0.260,)]TJ -16.8172 -1.0901 TD 0 Tc 0 Tw (F)Tj /F10 1 Tf 6.4 0 0 6.4 65.568 447.886 Tm -0.0019 Tc 0.0078 Tw (1, 18)Tj 2.0064 -0.6599 TD 0 Tc 0 Tw ( )Tj /F9 1 Tf 11 0 0 11 80.11 451.004 Tm -0.0057 Tc 0.0136 Tw (= 7.027, )Tj /F11 1 Tf 3.5459 0 TD 0 Tc 0 Tw (P)Tj /F9 1 Tf 0.603 0 TD -0.0056 Tc 0.0159 Tw [( = 0.015\).)]TJ /F12 1 Tf 1.9018 -2.1827 TD 0.0008 Tc (Discussion)Tj /F9 1 Tf -7.1021 -2.1801 TD -0.0055 Tc -0.0546 Tw [(The results of this study strongly support the view)]TJ -0.822 -1.0926 TD 0.0129 Tc 0.1227 Tw [(put forward by )49(Aebischer & Blake \(1994\), who)]TJ 0 -1.09 TD -0.0057 Tc 0.0785 Tw (suggested that grass strips, and in particular beetle)Tj T* -0.0061 Tc 0.1157 Tw (banks, may be valuable for game birds as well as)Tj 0 -1.0926 TD -0.0129 Tc -0.0609 Tw [(properly managed hedgerows. )43.8(Although beetle banks)]TJ 0 -1.09 TD -0.0059 Tc -0.0302 Tw (are simple landscape features, it was found that they)Tj T* 0.0395 Tc 0.1228 Tw (could develop high densities of the preferred)Tj 0 -1.0926 TD 0.0273 Tc 0.1227 Tw (invertebrate prey of game chicks, approaching)Tj 0 -1.09 TD 0.0532 Tc (equivalent values to those of more complex)Tj T* 0.0037 Tc (established hedgerows and other permanent field)Tj 0 -1.0926 TD 0.0196 Tc (boundaries. Despite overall chickfood densities)Tj 0 -1.09 TD -0.0059 Tc 0.0835 Tw (being lower in beetle banks, the numbers of many)Tj T* -0.0089 Tc -0.061 Tw [(key groups sampled often did not dif)11.7(fer significantly)60.7(.)]TJ 0 -1.0926 TD -0.0058 Tc -0.0553 Tw (This was particularly so for invertebrates considered)Tj 0 -1.09 TD -0.0037 Tc 0.1227 Tw (nutritionally important, such as sawfly larvae and)Tj T* -0.0104 Tc -0.061 Tw (caterpillars, found by Moreby \(1988\) to be especially)Tj 0 -1.0926 TD -0.0065 Tc 0.0569 Tw [(numerous in chick faecal samples.)]TJ /F12 1 Tf 13.7635 0 TD 0 Tc 0 Tw ( )Tj /F9 1 Tf 0.3015 0 TD -0.0066 Tc 0.0591 Tw (Faecal analysis,)Tj -14.065 -1.0901 TD 0.0063 Tc 0.1228 Tw [(although known to under)14(-represent softer)16.6(-bodied)]TJ T* -0.0084 Tc -0.061 Tw [(species, also found plant bugs, plant hoppers, ground)]TJ 0 -1.0926 TD -0.0057 Tc 0.1227 Tw [(beetles, leaf beetles, and weevils to be important)]TJ 0 -1.09 TD 0.0074 Tc [(dietary items. )10(These groups were also similar in)]TJ T* 0.0202 Tc (relative abundance within beetle banks and the)Tj 0 -1.0926 TD -0.0063 Tc 0.0601 Tw [(conventional field mar)11.7(gins sampled here.)]TJ 0.822 -1.09 TD -0.0059 Tc -0.0039 Tw (Invertebrate catch from the beetle banks and field)Tj -0.822 -1.09 TD 0.121 Tw [(m)7.2(a)7.2(r)12.2(gins varied considerably between the dif)12.2(ferent)]TJ 0 -1.0926 TD 0.0195 Tc 0.1227 Tw (fields, and also between farms. It is likely that)Tj 0 -1.0901 TD -0.0061 Tc -0.0418 Tw [(previous field management, including dif)9.4(ferences in)]TJ T* -0.0058 Tc -0.0071 Tw (cropping and agrochemical inputs over a number of)Tj 22.422 62.1792 TD -0.0057 Tc 0.0189 Tw (years, influenced catch. Increased replication in the)Tj 0 -1.09 TD -0.006 Tc -0.0197 Tw (experimental design would be necessary to evaluate)Tj T* 0.0068 Tc 0.1227 Tw (such factors, although complications such as the)Tj 0 -1.0926 TD -0.0056 Tc -0.0205 Tw [(frequent growing of dif)9.9(ferent crops on either side of)]TJ 0 -1.09 TD 0.0307 Tw (beetle banks, or the presence of other features such)Tj T* -0.006 Tc -0.0448 Tw [(as woods and roads adjacent to mar)12(gins, would have)]TJ 0 -1.0926 TD -0.0057 Tc 0.0374 Tw [(to be taken into account. In the first sampling year)33(,)]TJ 0 -1.0901 TD -0.0064 Tc -0.0453 Tw [(the farm used had well maintained field mar)11.6(gins and)]TJ T* -0.0057 Tc 0.1091 Tw (hedgerows following a history of environmentally)Tj 0 -1.0926 TD 0.0549 Tc 0.2295 Tw (conscious management to encourage game,)Tj 0 -1.09 TD -0.0055 Tc 0.0129 Tw (including the use of conservation headlands around)Tj T* 0.0163 Tc 0.1225 Tw [(all mar)8.6(gins. )13.5(This may explain why invertebrate)]TJ 0 -1.0926 TD 0.0139 Tc 0.1227 Tw (densities were similar in the habitats examined.)Tj 0 -1.0901 TD -0.0064 Tc 0.0006 Tw [(However in the second year)32.3(, when more farms with)]TJ T* 0.0183 Tc 0.1228 Tw [(dif)10.6(ferent management histories were compared,)]TJ 0 -1.0926 TD -0.0069 Tc -0.061 Tw (there was a greater variability in chickfood densities.)Tj 0 -1.09 TD 0.0003 Tc 0.1227 Tw (Barker & Reynolds \(1999\) also found significant)Tj T* -0.0061 Tc -0.0309 Tw (variation between farms when examining chickfood)Tj 0 -1.0926 TD -0.0063 Tc 0.0515 Tw [(abundance in grass mar)11.7(gins.)]TJ 0.8246 -1.09 TD 0.0008 Tc 0.1229 Tw [(Suctioning has been described as an inef)13.7(ficient)]TJ -0.8246 -1.09 TD 0.0101 Tc 0.1227 Tw (sampling method \(Green, 1984\), as chrysomelid)Tj 0 -1.0926 TD 0.0242 Tc (beetles, sawfly and lepidopteran larvae, highly)Tj 0 -1.09 TD -0.0059 Tc 0.0558 Tw (preferred dietary choices, may be scarce in suction)Tj T* -0.0128 Tc -0.061 Tw (samples though common in sweep net samples. More)Tj T* 0.027 Tc 0.1228 Tw [(recently)57.9(, sweep-netting has been preferentially)]TJ 0 -1.0926 TD -0.0058 Tc 0.0167 Tw (selected technique for chickfood capture \(Barker &)Tj 0 -1.0901 TD -0.0061 Tc 0.0242 Tw (Reynolds, 1999; Itamies )Tj /F11 1 Tf 9.9316 0 TD 0.0227 Tw (et al.)Tj /F9 1 Tf 1.992 0 TD -0.0057 Tc 0.0257 Tw (, 1996\). Beetle banks)Tj -11.9236 -1.09 TD 0.0224 Tc 0.1227 Tw (are established to enhance ground-active fauna)Tj 0 -1.0926 TD 0.0085 Tc (\(Thomas)Tj /F11 1 Tf 3.6232 0 TD 0.0106 Tc [( et al.)]TJ /F9 1 Tf 2.577 0 TD 0.0118 Tc 0.1228 Tw (, 1991, 1992\), for which suction-)Tj -6.2001 -1.09 TD 0.0067 Tc 0.1227 Tw (sampling may be a more appropriate assessment)Tj T* -0.0022 Tc (method.)Tj 0.8246 -1.0926 TD -0.0057 Tc 0.0602 Tw (Peak partridge chick hatch occurs in late June to)Tj -0.8246 -1.09 TD 0.1068 Tw [(early July)56.1(, coinciding with high insect abundance,)]TJ T* -0.007 Tc -0.0611 Tw [(although it may be any time from )44.6(April to September)]TJ 0 -1.0926 TD -0.0057 Tc 0.0843 Tw [(\(Green, 1984; )48.3(Anon., 1995\). )48.3(Although there was a)]TJ 0 -1.09 TD 0.0355 Tc 0.1227 Tw (lower catch of chickfood in beetle banks, the)Tj T* -0.0061 Tc 0.0759 Tw [(dif)11.9(ference between the habitats was consistent and)]TJ 0 -1.0926 TD -0.0058 Tc 0.001 Tw [(small in both May and )48.2(August, i.e. food availability)]TJ 0 -1.09 TD 0.0242 Tw (may remain adequate for chick survival throughout)Tj T* 0.0244 Tw (the hatching season. Chicks may fare better later in)Tj 0 -1.0926 TD -0.0057 Tc 0.1117 Tw (the season, when there is greater food abundance,)Tj 0 -1.09 TD -0.0061 Tc -0.0609 Tw [(with conventional field mar)9.4(gins providing improved)]TJ T* 0.0146 Tc 0.1227 Tw [(food resources. However)37.8(, the addition of beetle)]TJ 0 -1.0926 TD -0.0108 Tc -0.0613 Tw (banks to a monoculture of cereals may enhance game)Tj 0 -1.09 TD -0.0057 Tc -0.0455 Tw (chick survival, especially considering the increasing)Tj T* -0.0296 Tw (invertebrate diversity that appears to develop within)Tj 0 -1.0926 TD 0.0081 Tc 0.1227 Tw (the maturing habitat structure. Once established,)Tj 0 -1.0901 TD -0.0058 Tc 0.0443 Tw (beetle banks have a dense grassy structure, and are)Tj T* -0.0047 Tc 0.1228 Tw [(probably at least as good as many other mar)8.2(ginal)]TJ 0 -1.0926 TD -0.0118 Tc -0.061 Tw (habitats for provision of camouflaged shelter for both)Tj 0 -1.09 TD -0.0058 Tc -0.0398 Tw (adult game birds and their chicks, though may never)Tj T* 0.0106 Tc 0.1227 Tw (be equivalent to that provided by well managed)Tj 0 -1.0926 TD -0.011 Tc -0.0611 Tw (hedgerows. Plant cover was high throughout all sites,)Tj 0 -1.09 TD -0.0055 Tc 0.02 Tw [(and beetle banks were found to be valuable for the)]TJ T* -0.0058 Tc 0.0648 Tw (supply of nesting material, in late winter and early)Tj 0 -1.0926 TD 0.0028 Tc 0.1226 Tw [(spring. Many field mar)10.5(gins are observed to have)]TJ 0 -1.0901 TD -0.0059 Tc -0.0432 Tw (exposed bases, with little material of value for either)Tj T* 0.031 Tc 0.1227 Tw (forming a nest or allowing shelter from harsh)Tj ET endstream endobj 37 0 obj << /ProcSet [/PDF /Text ] /Font << /F4 4 0 R /F5 5 0 R /F6 6 0 R /F9 9 0 R /F10 10 0 R /F11 11 0 R /F12 12 0 R >> /ExtGState << /GS2 13 0 R >> >> endobj 39 0 obj << /Length 14462 >> stream BT /F4 1 Tf 9 0 0 9 522.205 788.95 Tm 0 0 0 1 k /GS2 gs -0.0019 Tc (117)Tj /F5 1 Tf 8 0 0 8 230.038 788.95 Tm -0.0057 Tc 0.0954 Tw [(Resour)29.7(ce pr)29.7(ovision for farmland gamebir)33.3(d)8.3(s)]TJ /F9 1 Tf 11 0 0 11 65.147 762.985 Tm 0.0411 Tc 0.1227 Tw (weather and predators, and so any additional)Tj 0 -1.09 TD -0.0075 Tc -0.0614 Tw [(resources in fields may be important. )10.5(The farms used)]TJ T* 0.0553 Tc 0.4432 Tw [(in this study tended to manage mar)14.1(gins)]TJ 0 -1.0926 TD -0.0058 Tc 0.1195 Tw [(sympathetically for wildlife and thus usually had)]TJ 0 -1.09 TD -0.0454 Tw (fairly well maintained hedgebases, inevitably linked)Tj T* 0.0422 Tw (to the desire to also create beetle banks.)Tj 0.822 -1.0926 TD -0.0057 Tc 0.0227 Tw (Foraging gamebirds tend to avoid vegetation that)Tj -0.822 -1.0901 TD 0.0227 Tc 0.1227 Tw [(is dif)12.4(ficult to penetrate \(P)30.4( )19.2(Thompson, personal)]TJ T* -0.0032 Tc (communication\). It may be that some of the sites)Tj 0 -1.0926 TD 0.0555 Tc 0.3236 Tw (developed cover that actually became too)Tj 0 -1.09 TD -0.006 Tc 0.0169 Tw (impenetrable as the season progressed, a factor that)Tj T* -0.0123 Tc -0.0614 Tw (may merely inconvenience adult birds, yet jeopardise)Tj 0 -1.0926 TD -0.0057 Tc -0.0045 Tw (the survival of chicks.)Tj /F12 1 Tf 8.6921 0 TD 0 Tc 0 Tw ( )Tj /F9 1 Tf 0.2397 0 TD -0.006 Tc -0.0045 Tw (Where plants are too densely)Tj -8.9317 -1.0901 TD 0.0474 Tc 0.1227 Tw (spaced, chicks may become so wet from the)Tj T* -0.0056 Tc 0.0199 Tw [(vegetation that unless they can dry of)12.4(f quickly they)]TJ 0 -1.0926 TD 0.0331 Tc 0.1227 Tw (may chill and die. Barker & Reynolds \(1999\))Tj 0 -1.09 TD 0.0345 Tc [(considered that many planted grassy mar)13.9(gins,)]TJ T* 0.0142 Tc (including beetle banks, could be less than ideal)Tj 0 -1.0926 TD -0.0056 Tc 0.0838 Tw (habitat for birds, although they reported that some)Tj 0 -1.09 TD -0.0076 Tc -0.0613 Tw [(farmers had experimentally cut channels within such)]TJ T* 0.0288 Tc 0.1227 Tw (habitats for birds to move around and dry out)Tj 0 -1.0926 TD 0.0263 Tc 0.1226 Tw [(following rain. )13.3(The provision of a sterile strip)]TJ 0 -1.09 TD 0.0173 Tc 0.1227 Tw (alongside field boundaries and features such as)Tj T* -0.0059 Tc 0.0198 Tw [(beetle banks has been suggested as useful for game)]TJ T* -0.0049 Tc 0.1227 Tw (chicks, as it provides an open area for drying out)Tj 0 -1.0926 TD -0.0093 Tc -0.061 Tw [(\(Bond, 1987\). Such solutions may be essential where)]TJ 0 -1.0901 TD 0.0507 Tc 0.1228 Tw [(vegetation has become especially dense. )12.1(W)87(e)]TJ T* -0.0056 Tc 0.05 Tw (observed that where tussocky cover had developed)Tj 0 -1.0926 TD -0.006 Tc 0.0234 Tw [(patchily)58.4(, often because many beetle banks are hand)]TJ 0 -1.09 TD -0.0058 Tc 0.0028 Tw (sown \(Thomas, 2000\), not only did this allow some)Tj T* -0.0057 Tc 0.0289 Tw (other plants to develop, but it allowed some degree)Tj 0 -1.0926 TD -0.006 Tc 0.0756 Tw [(of permeability for the benefit of game. )11.9(There is a)]TJ 0 -1.09 TD 0.0938 Tw [(clear trade-of)12(f between managing beetle banks for)]TJ T* 0.0316 Tc 0.1227 Tw (the dense tussock cover to enhance maximum)Tj 0 -1.0926 TD -0.0059 Tc 0.1155 Tw (predatory invertebrate survival, their primary aim,)Tj 0 -1.09 TD -0.0008 Tc 0.1227 Tw (and allowing some patchy alternative plant cover)Tj T* -0.0061 Tc 0.0418 Tw (for the benefit of game species.)Tj 0.822 -1.0926 TD 0.0335 Tw (Herbaceous and grass species that may be fed on)Tj -0.822 -1.09 TD 0.0013 Tc 0.1227 Tw (by adult birds were only present at low levels in)Tj T* 0.0341 Tc (beetle banks, although this has been found to)Tj 0 -1.0926 TD 0.0298 Tc 0.1228 Tw (increase through time following establishment)Tj 0 -1.09 TD 0.0428 Tc (\(Thomas )Tj /F11 1 Tf 4.2777 0 TD 0.0455 Tc 0.1227 Tw (et al.)Tj /F9 1 Tf 2.3991 0 TD (, 2000\). Many seed-producing)Tj -6.6769 -1.09 TD -0.0055 Tc 0.0084 Tw [(dicotyledons known to be preferred specifically by)]TJ 0 -1.0926 TD -0.0059 Tc 0.0466 Tw (game birds are more usually found within the field)Tj 0 -1.09 TD 0.0006 Tc 0.1227 Tw [(itself, rather than in boundary habitats. However)36.7(,)]TJ T* -0.0062 Tc 0.112 Tw [(with the development of more ef)11.8(ficient herbicides)]TJ 0 -1.0926 TD -0.0059 Tc -0.0449 Tw (and low tolerance of weeds within fields by farmers,)Tj 0 -1.0901 TD -0.0122 Tc -0.061 Tw [(weed seed availability may be low)60(, and thus any extra)]TJ T* -0.0091 Tc -0.0613 Tw (food resources within the habitat in which adult birds)Tj 0 -1.0926 TD -0.0059 Tc 0.0346 Tw (may be nesting can only be seen as beneficial.)Tj 0.822 -1.09 TD 0.0112 Tc 0.1227 Tw (Game birds with chicks were observed within)Tj -0.822 -1.09 TD -0.0055 Tc -0.0056 Tw [(banks during the course of the study)58.9(, indicating that)]TJ 0 -1.0926 TD -0.0058 Tc 0.0812 Tw (the habitat was being utilised. Overall, we suggest)Tj 0 -1.09 TD -0.0028 Tc 0.1227 Tw (that beetle banks make a valuable contribution to)Tj T* -0.006 Tc 0.0915 Tw (game habitat on farmland, with their low cost and)Tj 0 -1.0926 TD -0.0055 Tc 0.0561 Tw (ease of construction adding further positive points.)Tj 0 -1.0901 TD -0.0061 Tc 0.0242 Tw (They may be especially invaluable when combined)Tj T* -0.0008 Tc 0.1228 Tw (with spring brood-rearing and winter cover crops)Tj 22.422 62.1792 TD 0.007 Tc 0.1227 Tw [(within set-aside strips, such as recommended by)]TJ 0 -1.09 TD -0.006 Tc -0.0052 Tw (Boatman & Bence \(2000\), a methodology shown to)Tj T* -0.0058 Tc 0.02 Tw (significantly increase wild pheasant populations, as)Tj 0 -1.0926 TD 0.0558 Tc 0.1273 Tw (well as the nationally declining skylark, on)Tj 0 -1.09 TD -0.006 Tc 0.06 Tw (conventionally managed farmland on UK lowland.)Tj T* 0.0556 Tc 0.2664 Tw (Although beetle banks may add chickfood)Tj 0 -1.0926 TD -0.0057 Tc -0.0545 Tw (invertebrates and nesting shelter to arable fields, and)Tj 0 -1.0901 TD 0.0039 Tc 0.1227 Tw (may be useful where such resources are lacking,)Tj T* -0.0058 Tc -0.0209 Tw (appropriate good management of conventional field)Tj 0 -1.0926 TD 0.0325 Tc 0.1227 Tw [(m)6.9(a)6.9(r)9.3(gins may be much more important for the)]TJ 0 -1.09 TD 0.0506 Tc (continuing survival of game bird species on)Tj T* -0.0004 Tc (farmland.)Tj /F12 1 Tf 6.0275 -2.1827 TD 0.002 Tc (Acknowledgements)Tj /F9 1 Tf -5.2029 -2.1827 TD 0.0065 Tc (The authors wish to thank R Noordhuis and S)Tj -0.8246 -1.09 TD -0.0022 Tc [(Campbell for fieldwork assistance, and H Oliver)10.7(-)]TJ T* -0.0062 Tc -0.0472 Tw [(Bellasis, R S)11.8(tirling, R Ody)58.2(, G Blatchford, D Bowyer)]TJ 0 -1.0926 TD -0.0018 Tc 0.1228 Tw (and M Malyon for generously allowing access to)Tj 0 -1.09 TD -0.006 Tc -0.0467 Tw [(field sites. )48.2(Anonymous referees are thanked for their)]TJ T* -0.0062 Tc 0.0202 Tw [(useful criticism of an earlier draft. )11.9(The Leverhulme)]TJ 0 -1.0926 TD -0.0052 Tc 0.0659 Tw [(T)36.1(rust provided funding.)]TJ /F12 1 Tf 7.8468 -2.1801 TD 0.0015 Tc [(Refer)6.7(ences)]TJ /F19 1 Tf 9 0 0 9 311.789 467.36 Tm -0.0041 Tc -0.0057 Tw [(Aebischer)11.6( N J, Blake K )50.2(A. 1994.)]TJ /F4 1 Tf 13.4551 0 TD -0.0055 Tc -0.0078 Tw [( Field mar)10.3(gins as habitats for)]TJ -12.4473 -1.1118 TD -0.0064 Tc -0.0555 Tw (game. In )Tj /F20 1 Tf 3.5874 0 TD -0.0052 Tc -0.0578 Tw [(British Cr)32.6(op Pr)35.7(otection Council Monograph No.58:)]TJ -3.5874 -1.1118 TD -0.0055 Tc -0.0241 Tw [(Field mar)35.4(gins: Integrating agricultur)32.3(e and conser)-5.5(vation)]TJ /F4 1 Tf 22.1606 0 TD -0.0028 Tc -0.0277 Tw (, pp.)Tj -22.1606 -1.1118 TD -0.0052 Tc 0.06 Tw (95-104. Farnham: British Crop Protection Council.)Tj /F19 1 Tf -1.0079 -1.1087 TD 0.004 Tc 0.1223 Tw (Anon. 1995.)Tj /F4 1 Tf 5.2504 0 TD 0 Tc 0 Tw ( )Tj /F20 1 Tf 0.3843 0 TD 0.003 Tc 0.1264 Tw [(Farmland bir)31.3(d)-0.1( management guidelines leaflet)]TJ /F4 1 Tf 18.9921 0 TD 0 Tc 0 Tw (.)Tj -23.6189 -1.1118 TD -0.0042 Tc 0.0199 Tw (Sandy: Royal Society for the Protection of Birds.)Tj /F19 1 Tf -1.0079 -1.1118 TD -0.0087 Tc -0.0611 Tw (Anon. 1999)Tj /F21 1 Tf 4.6898 0 TD 0 Tc 0 Tw (a)Tj /F19 1 Tf 0.4882 0 TD (.)Tj /F4 1 Tf 0.252 0 TD ( )Tj /F20 1 Tf 0.1827 0 TD -0.0097 Tc -0.0735 Tw [(The Arable )-12.7(S)28.1(tewar)31.2(dship )-12.3(Scheme:)]TJ /F4 1 Tf 12.7591 0 TD 0 Tc 0 Tw ( )Tj /F20 1 Tf 0.1858 0 TD -0.0102 Tc -0.0667 Tw (Information and)Tj -17.5496 -1.1118 TD -0.005 Tc -0.0333 Tw [(how to apply)54.8(.)-6.4( )]TJ /F4 1 Tf 5.4551 0 TD -0.0048 Tc -0.0347 Tw (PB3257A. London: MAFF Publications. 56pp.)Tj /F19 1 Tf -6.463 -1.1118 TD 0.0576 Tc 0.1833 Tw (Anon. 1999)Tj /F21 1 Tf 5.5937 0 TD 0 Tc 0 Tw (b)Tj /F19 1 Tf 0.5575 0 TD (.)Tj /F4 1 Tf 0.315 0 TD ( )Tj /F20 1 Tf 0.5008 0 TD 0.0571 Tc 0.1867 Tw [(The Countr)-12.2(yside S)31.9(tewar)35.1(dship Scheme:)]TJ -5.9591 -1.1087 TD -0.0041 Tc 0.1244 Tw [(Information and how to apply)52.6(.)-5.2( )]TJ /F4 1 Tf 12.8976 0 TD 0.1278 Tw (PB3950A. London: MAFF)Tj -12.8976 -1.1118 TD -0.0046 Tc 0.1277 Tw (Publications. 80pp.)Tj /F19 1 Tf -1.0079 -1.1118 TD -0.0038 Tc -0.037 Tw (Asteraki E J, Hanks C B, Clements R O. 1992.)Tj /F4 1 Tf 19.2283 0 TD -0.0068 Tc [( )9.8(The impact of)]TJ -18.2205 -1.1118 TD 0.0418 Tc 0.125 Tw (the chemical removal of the hedge-base flora on the)Tj 0 -1.1087 TD 0.0482 Tc 0.1255 Tw (community structure of carabid beetles \(Coleoptera,)Tj 0 -1.1118 TD 0.0089 Tc 0.125 Tw (Carabidae\) and spiders \(Araneae\) of the field and hedge)Tj T* -0.007 Tc (bottom. )Tj /F20 1 Tf 3.348 0 TD -0.0054 Tc 0.0686 Tw [(Journal of )10.4(Applied Entomology)]TJ /F4 1 Tf 12.6268 0 TD 0 Tc 0 Tw ( )Tj /F19 1 Tf 0.3244 0 TD -0.0632 Tc [(11)-50.2(3)]TJ /F4 1 Tf 1.4268 0 TD -0.0061 Tc (:398-406.)Tj /F19 1 Tf -18.7339 -1.1118 TD -0.0038 Tc -0.0333 Tw (Asteraki E J, Hanks C B, Clements R O. 1995.)Tj /F4 1 Tf 19.263 0 TD -0.0058 Tc -0.0511 Tw [( The )-14.9(influence)]TJ -18.2551 -1.1118 TD -0.0057 Tc -0.0019 Tw [(of dif)10(ferent types of grassland field mar)10.1(gin on carabid beetle)]TJ 0 -1.1087 TD 0.0568 Tc 0.2648 Tw (\(Coleoptera, Carabidae\) communities. )Tj /F20 1 Tf 18.4221 0 TD 0.0563 Tc [(Agricultur)28(e,)]TJ -18.4221 -1.1118 TD -0.0054 Tc 0.0728 Tw [(Ecosystems and Envir)35.5(onment)]TJ /F4 1 Tf 11.748 0 TD 0 Tc 0 Tw ( )Tj /F19 1 Tf 0.3213 0 TD -0.013 Tc (54)Tj /F4 1 Tf 0.989 0 TD -0.0061 Tc (:195-202.)Tj /F19 1 Tf -14.0661 -1.1118 TD -0.0043 Tc -0.0033 Tw [(Bannister)11.4( N R, )13.7(W)62(att )13.7(T)17.7( )52.5(A. 1994.)]TJ /F4 1 Tf 13.0835 0 TD -0.0063 Tc -0.0055 Tw [( Hedgerow management: Past)]TJ -12.0756 -1.1118 TD 0.0578 Tc 0.1741 Tw (and present. In )Tj /F20 1 Tf 7.6157 0 TD 0.0572 Tc 0.1762 Tw [(Hedger)32(ow management and natur)32(e)]TJ -7.6157 -1.1087 TD 0.0101 Tc (conservation)Tj /F4 1 Tf 5.2913 0 TD 0.0127 Tc 0.1248 Tw [(, pp. 7-15. Eds )12.5(T)15.7( )52.7(A)56.7( )8.6(W)88.5(att and G P)34.7( Buckley)63.1(.)]TJ -5.2913 -1.1118 TD -0.0044 Tc 0.0266 Tw [(London: )15.2(W)71.4(ye College Press.)]TJ /F19 1 Tf -1.0079 -1.1118 TD -0.0031 Tc 0.1252 Tw [(Barker)12.7( )53(A)56.7( M, Reynolds C J M. 1999.)]TJ /F4 1 Tf 15.9685 0 TD -0.0042 Tc 0.1264 Tw [( )12.2(The value of planted)]TJ -14.9606 -1.1118 TD -0.0052 Tc 0.0156 Tw [(grass field mar)13.7(gins as a habitat for sawflies and other chick-)]TJ 0 -1.1087 TD -0.0048 Tc 0.0444 Tw (food insects. )Tj /F20 1 Tf 5.326 0 TD -0.0053 Tc [(Aspects of )10.5(Applied Biology)]TJ /F4 1 Tf 10.7937 0 TD 0 Tc 0 Tw ( )Tj /F19 1 Tf 0.3024 0 TD -0.013 Tc (54)Tj /F4 1 Tf 0.9858 0 TD -0.0074 Tc [(:109-1)27.2(16.)]TJ /F19 1 Tf -18.4157 -1.1118 TD -0.004 Tc 0.0606 Tw [(Batten L)52.7( )50.5(A, Bibby C J, Clement P)84.2(,)0.1( Elliot G)33.8( D, Porter)14.9( R F)84.2(.)]TJ 1.0079 -1.1118 TD -0.0065 Tc (1990.)Tj /F4 1 Tf 2.2268 0 TD 0 Tc 0 Tw ( )Tj /F20 1 Tf 0.2835 0 TD -0.0049 Tc 0.0361 Tw [(Red data bir)36(ds in Britain)]TJ /F4 1 Tf 9.9874 0 TD -0.0041 Tc 0.0389 Tw [(. London: Poyser)52.6(.)]TJ /F19 1 Tf -13.5055 -1.1118 TD -0.0043 Tc -0.0007 Tw [(Bence S, S)46.1(tander)11.5( K, Griffiths M. 1999. )]TJ /F4 1 Tf 16.4504 0 TD -0.0041 Tc -0.0018 Tw (Nest site selection by)Tj -15.4425 -1.1118 TD 0.1259 Tw (the harvest mouse )Tj /F20 1 Tf 7.7827 0 TD -0.004 Tc 0.123 Tw [(Micr)33.8(opuys minutus)]TJ /F4 1 Tf 7.7165 0 TD -0.0036 Tc 0.1277 Tw [( on arable farmland.)]TJ /F20 1 Tf -15.4992 -1.1087 TD -0.0052 Tc 0.0593 Tw [(Aspects of )13.7(Applied Biology)]TJ /F4 1 Tf 10.8378 0 TD 0 Tc 0 Tw ( )Tj /F19 1 Tf 0.3118 0 TD -0.013 Tc (54)Tj /F4 1 Tf 0.989 0 TD -0.0061 Tc (:197-202.)Tj /F19 1 Tf -13.1465 -1.1118 TD -0.0037 Tc 0.0454 Tw (Boatman N D, Bence S L. 2000.)Tj /F4 1 Tf 13.5181 0 TD -0.0059 Tc 0.0459 Tw [( Management of set-aside to)]TJ -12.5102 -1.1118 TD -0.0046 Tc 0.0421 Tw (enhance biodiversity: the wild bird cover option. )Tj /F20 1 Tf 19.7921 0 TD -0.005 Tc 0.0444 Tw (Aspects of)Tj -19.7921 -1.1118 TD -0.0058 Tc 0.0889 Tw (Applied Biology)Tj /F4 1 Tf 6.4819 0 TD 0 Tc 0 Tw ( )Tj /F19 1 Tf 0.3402 0 TD -0.013 Tc (62)Tj /F4 1 Tf 0.989 0 TD -0.006 Tc (:73-78.)Tj /F19 1 Tf -8.8189 -1.1087 TD -0.0091 Tc -0.0611 Tw (Bond S D. 1987.)Tj /F4 1 Tf 6.485 0 TD -0.0122 Tc -0.0635 Tw [( Field mar)9.8(gins, a farmer)-46.8(\222)60.4(s)-0.2( view on management.)]TJ -5.4772 -1.1118 TD -0.0064 Tc (In )Tj /F20 1 Tf 1.0268 0 TD -0.005 Tc -0.0444 Tw [(British Cr)35.9(op Pr)32.8(otection Council Monograph No. 35:)]TJ /F4 1 Tf 20.5543 0 TD 0 Tc 0 Tw ( )Tj /F20 1 Tf 0.2047 0 TD -0.0072 Tc (Field)Tj -21.7858 -1.1118 TD -0.0007 Tc [(mar)33.9(gins)]TJ /F4 1 Tf 3.2441 0 TD 0.0029 Tc 0.1255 Tw [(, pp. 79-83. Eds J M )16(W)88.1(ay and P)37.5( )12.1(W)28.3( Greig-Smith.)]TJ -3.2441 -1.1118 TD -0.0052 Tc 0.0597 Tw [(Farnham: British Crop Protection Council.)]TJ /F19 1 Tf -1.0079 -1.1118 TD -0.0039 Tc 0.125 Tw [(Chamberlain D E, Fuller R J. 2000.)]TJ /F4 1 Tf 15.7449 0 TD -0.005 Tc 0.126 Tw [( Local extinctions and)]TJ ET endstream endobj 40 0 obj << /ProcSet [/PDF /Text ] /Font << /F4 4 0 R /F5 5 0 R /F9 9 0 R /F11 11 0 R /F12 12 0 R /F19 41 0 R /F20 42 0 R /F21 43 0 R >> /ExtGState << /GS2 13 0 R >> >> endobj 45 0 obj << /Length 16632 >> stream BT /F4 1 Tf 9 0 0 9 59.502 788.95 Tm 0 0 0 1 k /GS2 gs -0.0019 Tc (118)Tj /F6 1 Tf 8 0 0 8 258.948 788.95 Tm -0.0084 Tc 0.0207 Tw [(S R )11.3(THOMAS )]TJ /F5 1 Tf 6.1618 0 TD -0.0051 Tc 0.0052 Tw [(E)5.3(T)12.6( AL.)]TJ /F4 1 Tf 9 0 0 9 68.545 764.346 Tm 0.0048 Tc 0.125 Tw (changes in species richness of lowland farmland birds in)Tj 0 -1.1118 TD -0.0107 Tc -0.0623 Tw [(England and )11.4(W)87.1(ales in relation to recent changes in agricultural)]TJ T* -0.0048 Tc (land-use. )Tj /F20 1 Tf 3.8173 0 TD -0.0052 Tc -0.0326 Tw [(Agricultur)32.6(e, Ecosystems and Envir)32.6(onment)]TJ /F4 1 Tf 16.5165 0 TD 0 Tc 0 Tw ( )Tj /F19 1 Tf 0.2173 0 TD -0.013 Tc (78)Tj /F4 1 Tf 0.989 0 TD -0.0059 Tc (:1-17.)Tj /F19 1 Tf -22.5449 -1.1087 TD 0.0088 Tc 0.125 Tw [(Collins K L, )15.1(W)24.7(i)0.7(lcox )52.9(A, Chaney K, Boatman N D. 1996.)]TJ /F4 1 Tf 1.0047 -1.1118 TD 0.0158 Tc 0.1256 Tw (Relationship between polyphagous predator density and)Tj T* -0.0055 Tc 0.0333 Tw [(overwintering habitat within arable field mar)7.1(gins and beetle)]TJ T* -0.0058 Tc (banks.)Tj /F20 1 Tf 2.5575 0 TD -0.0046 Tc 0.0215 Tw [( The 1996 Brighton Confer)36.3(ence - Pests and Diseases)]TJ /F4 1 Tf 21.0646 0 TD 0 Tc 0 Tw (,)Tj -23.6221 -1.1087 TD -0.0051 Tc 0.0945 Tw (pp. 635-640.)Tj /F19 1 Tf -1.0047 -1.1118 TD 0.034 Tc 0.1259 Tw [(Dunkley F)30.9( )56.2(A. 1997.)]TJ /F4 1 Tf 9.0961 0 TD 0 Tc 0 Tw ( )Tj /F20 1 Tf 0.4095 0 TD 0.0333 Tc 0.1278 Tw [(Management options for hedger)30.1(o)5.1(w)]TJ -8.5008 -1.1118 TD 0.0574 Tc 0.3144 Tw [(vegetation: Combining weed contr)32.2(ol with habitat)]TJ T* 0.0026 Tc 0.1252 Tw [(impr)34.1(ovement for pr)27.8(edatory ar)-10(thr)37.2(opods)]TJ /F4 1 Tf 15.9118 0 TD 0.0044 Tc 0.1277 Tw (. Unpublished PhD)Tj -15.9118 -1.1087 TD -0.0051 Tc 0.037 Tw (thesis. University of Southampton.)Tj /F19 1 Tf -1.0047 -1.1118 TD -0.0041 Tc -0.0277 Tw [(Ford J, Chitty H, Middleton )49.5(A)55.7( D. 1938.)]TJ /F4 1 Tf 16.2835 0 TD -0.0059 Tc [( )10.6(The food of partridge)]TJ -15.2787 -1.1118 TD 0.0057 Tc (chicks )Tj /F20 1 Tf 2.9732 0 TD 0.0051 Tc 0.1296 Tw [(Per)30.3(dix per)36.6(dix)]TJ /F4 1 Tf 5.6063 0 TD 0.0077 Tc 0.125 Tw [( in Great Britain. )]TJ /F20 1 Tf 7.6724 0 TD 0.0063 Tc 0.1282 Tw [(Journal of )13(Animal)]TJ -16.252 -1.1118 TD -0.007 Tc (Ecology)Tj /F4 1 Tf 3.2409 0 TD 0 Tc 0 Tw ( )Tj /F19 1 Tf 0.378 0 TD (7)Tj /F4 1 Tf 0.4945 0 TD -0.0053 Tc (:251-265.)Tj /F19 1 Tf -5.1181 -1.1118 TD -0.0047 Tc -0.0611 Tw [(G)11.2(r)11(een R E. 1984.)]TJ /F4 1 Tf 7.0236 0 TD -0.0063 Tc -0.0635 Tw [( )9.1(The feeding ecology and survival of partridge)]TJ -6.0189 -1.1087 TD -0.0051 Tc (chicks )Tj /F20 1 Tf 2.7402 0 TD -0.0056 Tc -0.0327 Tw (Alectoris rufa)Tj /F4 1 Tf 5.4236 0 TD -0.0042 Tc -0.0362 Tw [( and )]TJ /F20 1 Tf 1.8551 0 TD -0.0065 Tc -0.0376 Tw [(Per)31.3(dix per)34.4(dix)]TJ /F4 1 Tf 5.2882 0 TD -0.0059 Tc -0.0361 Tw [( on arable farmland in)]TJ -15.3071 -1.1118 TD -0.0045 Tc -0.0046 Tw (East Anglia. )Tj /F20 1 Tf 5.1748 0 TD -0.0054 Tc 0.0464 Tw [(Journal of )13.6(Applied Ecology)]TJ /F4 1 Tf 11.0205 0 TD 0 Tc 0 Tw ( )Tj /F19 1 Tf 0.2992 0 TD -0.013 Tc (21)Tj /F4 1 Tf 0.989 0 TD -0.0061 Tc (:817-830.)Tj /F19 1 Tf -18.4882 -1.1118 TD -0.0054 Tc -0.0621 Tw [(G)10.5(r)10.3(egory R D, Noble D G)114.3(,)0.5( Campbell L)48.1( H, Gibbons D )13.9(W)98.6(.)-0.9( 2000.)]TJ /F20 1 Tf 1.0047 -1.1118 TD -0.0048 Tc 0.063 Tw [(The state of the UK\222)121.2(s bir)33(ds 1999)]TJ /F4 1 Tf 13.1244 0 TD -0.0035 Tc 0.0639 Tw (. Sandy: Royal Society for)Tj -13.1244 -1.1087 TD -0.0048 Tc 0.0117 Tw [(the Protection of Birds and British )10.9(T)32.8(rust for Ornithology)64.5(. 23)]TJ 0 -1.1118 TD 0.0044 Tc (pp.)Tj /F19 1 Tf -1.0047 -1.1118 TD -0.0044 Tc 0.0444 Tw [(Hill D )49.1(A. 1985.)]TJ /F4 1 Tf 6.3213 0 TD -0.0051 Tc [( )8.1(The feeding ecology and survival of pheasant)]TJ -5.3165 -1.1118 TD 0.0212 Tc 0.125 Tw (chicks on arable farmland. )Tj /F20 1 Tf 11.9402 0 TD 0.0216 Tc 0.126 Tw [(Journal of )15.4(Applied Ecology)]TJ /F19 1 Tf -11.9402 -1.1118 TD -0.0067 Tc (22)Tj /F4 1 Tf 1.0079 0 TD 0.0041 Tc (:645-654.)Tj /F19 1 Tf -2.0126 -1.1087 TD 0.0203 Tc 0.125 Tw [(Hill D )45.5(A, Robertson P)51.8( )51(A. 1988.)]TJ /F4 1 Tf 14.2835 0 TD 0 Tc 0 Tw ( )Tj /F20 1 Tf 0.3937 0 TD 0.0197 Tc 0.1278 Tw [(The Pheasant: Ecology)54.3(,)]TJ -13.6724 -1.1118 TD -0.0085 Tc -0.0611 Tw (Management and Conservation)Tj /F4 1 Tf 12.2614 0 TD -0.063 Tw (. Oxford: Blackwell Scientific)Tj -12.2614 -1.1118 TD -0.0047 Tc 0.0722 Tw (Publications. 278 pp.)Tj /F19 1 Tf -1.0047 -1.1118 TD -0.0041 Tc 0.1223 Tw [(Itamies J, Putaala )52.7(A, Pirinen M, Hissa R. 1996.)]TJ /F4 1 Tf 20.8 0 TD -0.0063 Tc 0.1132 Tw [( The )-11.8(food)]TJ -19.7953 -1.1087 TD -0.0053 Tc -0.0234 Tw (composition of grey partridge chicks )Tj /F20 1 Tf 14.6677 0 TD -0.0066 Tc -0.021 Tw [(Per)31.2(dix per)31.2(dix)]TJ /F4 1 Tf 5.3071 0 TD -0.005 Tc -0.025 Tw [( in central)]TJ -19.9748 -1.1118 TD -0.0047 Tc (Finland. )Tj /F20 1 Tf 3.578 0 TD -0.0055 Tc 0.0612 Tw (Ornis Fennica)Tj /F4 1 Tf 5.8016 0 TD 0 Tc 0 Tw ( )Tj /F19 1 Tf 0.315 0 TD -0.013 Tc (73)Tj /F4 1 Tf 0.989 0 TD -0.006 Tc (:27-34.)Tj /F19 1 Tf -11.6882 -1.1118 TD -0.0038 Tc -0.0406 Tw (Luff M L. 1965.)Tj /F4 1 Tf 6.5669 0 TD -0.0063 Tc -0.0416 Tw [( )12.1(The morphology and microclimate of )]TJ /F20 1 Tf 15.0236 0 TD -0.006 Tc (Dactylis)Tj -20.5858 -1.1118 TD -0.0064 Tc (glomerata)Tj /F4 1 Tf 4.063 0 TD -0.004 Tc 0.0556 Tw [( tussocks. )]TJ /F20 1 Tf 4.2047 0 TD -0.0054 Tc 0.0584 Tw (Journal of Ecology)Tj /F4 1 Tf 7.6945 0 TD 0 Tc 0 Tw ( )Tj /F19 1 Tf 0.3118 0 TD -0.013 Tc (53)Tj /F4 1 Tf 0.989 0 TD -0.0061 Tc (:771-787.)Tj /F19 1 Tf -18.2677 -1.1118 TD -0.0043 Tc -0.0024 Tw [(MacLeod A. )-53.5(1994. )]TJ /F4 1 Tf 8.0504 0 TD -0.0054 Tc 0.05 Tw (Provision of plant resources for beneficial)Tj -7.0457 -1.1087 TD -0.0049 Tc 0.1009 Tw [(arthropods in arable ecosystems. Unpublished PhD )10.8(Thesis.)]TJ 0 -1.1118 TD -0.0056 Tc 0.0445 Tw (University of Southampton.)Tj /F19 1 Tf -1.0047 -1.1118 TD -0.0102 Tc -0.0598 Tw [(Magurran )51(A)49.6( E. 1988.)]TJ /F4 1 Tf 8.6047 0 TD 0 Tc 0 Tw ( )Tj /F20 1 Tf 0.1795 0 TD -0.0106 Tc -0.0623 Tw [(Ecological diversity and its measur)33.5(ement)]TJ /F4 1 Tf 15.8425 0 TD 0 Tc 0 Tw (.)Tj -23.6221 -1.1118 TD -0.0051 Tc 0.0367 Tw (London: Chapman and Hall. 179 pp.)Tj /F19 1 Tf -1.0047 -1.1087 TD -0.0043 Tc 0.0426 Tw [(Middleton )52.8(A)55.5( D, Chitty H. 1937.)]TJ /F4 1 Tf 13.3543 0 TD -0.0056 Tc 0.0422 Tw [( )5.3(The food of adult partridges)]TJ /F20 1 Tf -12.3496 -1.1118 TD -0.0065 Tc 0.0124 Tw [(Per)31.3(dix per)31.3(dix)]TJ /F4 1 Tf 5.3386 0 TD -0.0041 Tc 0.0139 Tw [( and )]TJ /F20 1 Tf 1.9559 0 TD -0.0061 Tc 0.0123 Tw [(Alectoris r)-9.2(ufa)]TJ /F4 1 Tf 5.4677 0 TD -0.004 Tc 0.0139 Tw [( in Great Britain. )]TJ /F20 1 Tf 7.0047 0 TD -0.0055 Tc 0.0166 Tw (Journal of)Tj -19.7669 -1.1118 TD -0.0058 Tc 0.0889 Tw (Applied Ecology)Tj /F4 1 Tf 6.652 0 TD 0 Tc 0 Tw ( )Tj /F19 1 Tf 0.3433 0 TD (6)Tj /F4 1 Tf 0.4945 0 TD -0.0061 Tc (:322-336.)Tj /F19 1 Tf -8.4945 -1.1118 TD -0.0034 Tc 0.1241 Tw [(M)6.3(o)6.3(r)15.5(eby S J. 1988.)]TJ /F4 1 Tf 7.9843 0 TD -0.0047 Tc 0.1262 Tw [( )49.3(An aid to the identification of arthropod)]TJ -6.9795 -1.1087 TD -0.0078 Tc -0.0625 Tw (fragments in the faeces of gamebird chicks \(Galliformes\). )Tj /F20 1 Tf 22.4063 0 TD -0.0106 Tc (Ibis)Tj /F19 1 Tf -22.4063 -1.1118 TD -0.0019 Tc (130)Tj /F4 1 Tf 1.5118 0 TD 0.0041 Tc (:519-526.)Tj /F19 1 Tf -2.5165 -1.1118 TD -0.0038 Tc 0.0193 Tw [(M)5.9(o)5.9(r)15.1(eby S J, )50.5(Aebischer)11.9( N J. 1992.)]TJ /F4 1 Tf 14.085 0 TD -0.0054 Tc 0.0203 Tw [( Invertebrate abundance on)]TJ -13.0803 -1.1118 TD -0.0134 Tc -0.0625 Tw (cereal fields and set-aside land: implications for wild gamebird)Tj T* -0.0042 Tc 0.0722 Tw (chicks. In )Tj /F20 1 Tf 4.2394 0 TD -0.0051 Tc 0.0754 Tw [(British Cr)32.7(op Pr)35.8(otection Council Monograph No.)]TJ -4.2394 -1.1087 TD -0.0059 Tc 0.1 Tw (50: Set-aside)Tj /F4 1 Tf 5.285 0 TD -0.0044 Tc 0.1048 Tw (, pp. 181-186. Ed. J Clarke. Farnham: British)Tj -5.285 -1.1118 TD -0.005 Tc 0.0806 Tw (Crop Protection Council.)Tj /F19 1 Tf 26.4 53.3323 TD -0.0035 Tc -0.0509 Tw [(M)6.2(o)6.2(r)12.3(eby S J, Southway S E. 1999.)]TJ /F4 1 Tf 13.6882 0 TD -0.0062 Tc -0.0514 Tw [( Influence of autumn applied)]TJ -12.6803 -1.1118 TD -0.0058 Tc 0.0191 Tw (herbicides on summer and autumn food available to birds in)Tj T* 0.0352 Tc 0.125 Tw (winter wheat fields in southern England. )Tj /F20 1 Tf 18.6646 0 TD 0.0342 Tc [(Agricultur)27.9(e,)]TJ -18.6646 -1.1087 TD -0.0054 Tc 0.0728 Tw [(Ecosystems and Envir)35.5(onment)]TJ /F4 1 Tf 11.748 0 TD 0 Tc 0 Tw ( )Tj /F19 1 Tf 0.3213 0 TD -0.013 Tc (72)Tj /F4 1 Tf 0.989 0 TD -0.0061 Tc (:285-297.)Tj /F19 1 Tf -14.0661 -1.1118 TD 0.0299 Tc 0.1253 Tw [(M)6.6(o)6.6(r)14.2(eby S J, Southway S E, Boatman N D. 1999.)]TJ /F4 1 Tf 22.8724 0 TD 0.0282 Tc 0.1165 Tw [( The)]TJ -21.8646 -1.1118 TD -0.0033 Tc 0.1253 Tw (importance of the crop edge compared to the mid-field in)Tj T* -0.0023 Tc 0.125 Tw (providing invertebrate food for farmland birds. )Tj /F20 1 Tf 19.6756 0 TD -0.0013 Tc 0.1277 Tw (Aspects of)Tj -19.6756 -1.1087 TD -0.0057 Tc 0.1 Tw (Applied Biology)Tj /F4 1 Tf 6.4945 0 TD 0 Tc 0 Tw ( )Tj /F19 1 Tf 0.3496 0 TD -0.013 Tc (54)Tj /F4 1 Tf 0.989 0 TD -0.0061 Tc (:217-222.)Tj /F19 1 Tf -8.8409 -1.1118 TD -0.0075 Tc -0.0611 Tw (Pollard E. 1968.)Tj /F4 1 Tf 6.589 0 TD -0.01 Tc -0.0624 Tw [( Hedges III. )5.7(The ef)5.7(fect of removal of the bottom)]TJ -5.5811 -1.1118 TD -0.0051 Tc 0.0044 Tw (flora of a hawthorn hedgerow on the Carabidae of the hedge)Tj T* -0.0072 Tc (bottom. )Tj /F20 1 Tf 3.3386 0 TD -0.0053 Tc 0.0592 Tw [(Journal of )13.5(Applied Ecology)]TJ /F4 1 Tf 11.0614 0 TD 0 Tc 0 Tw ( )Tj /F19 1 Tf 0.3087 0 TD (5)Tj /F4 1 Tf 0.4945 0 TD -0.0061 Tc (:125-139.)Tj /F19 1 Tf -16.211 -1.1087 TD 0.0043 Tc 0.1244 Tw [(Potts G)35.8( R. 1986.)]TJ /F4 1 Tf 7.3291 0 TD 0 Tc 0 Tw ( )Tj /F20 1 Tf 0.3874 0 TD 0.0042 Tc 0.1263 Tw [(The Par)-8.4(tridge: Pesticides, Pr)35.7(edation and)]TJ -6.7087 -1.1118 TD -0.0059 Tc (Conservation)Tj /F4 1 Tf 5.3197 0 TD -0.0042 Tc 0.0569 Tw (. London: Collins. 274 pp.)Tj /F19 1 Tf -6.3276 -1.1118 TD -0.0062 Tc -0.0611 Tw (Potts G R. 1997.)Tj /F4 1 Tf 6.6394 0 TD -0.0069 Tc -0.0629 Tw [( Cereal farming, pesticides and grey partridges.)]TJ -5.6315 -1.1118 TD -0.0052 Tc (In )Tj /F20 1 Tf 1.115 0 TD -0.005 Tc 0.0503 Tw [(Farming and Bir)35.9(ds in Eur)32.8(ope: the Common )14.2(Agricultural)]TJ -1.115 -1.1118 TD 0.0642 Tw [(Policy and its Implications for Bir)35.9(d Conser)-5(vation)]TJ /F4 1 Tf 19.937 0 TD -0.0038 Tc 0.0666 Tw (, pp. 150-)Tj -19.937 -1.1087 TD -0.0123 Tc -0.0619 Tw [(177. Eds D J Pain and M )13.6(W)28.8( Pienkowski. 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Tf 5.5118 0 TD 0.0031 Tc (. )Tj /F20 1 Tf 0.5512 0 TD -0.0056 Tc 0.0555 Tw (Ornis Scandinavica)Tj /F4 1 Tf 7.8772 0 TD 0 Tc 0 Tw ( )Tj /F19 1 Tf 0.3055 0 TD -0.013 Tc (19)Tj /F4 1 Tf 0.989 0 TD -0.006 Tc (:35-40.)Tj /F19 1 Tf -20.1008 -1.1118 TD -0.0601 Tw [(Sother)-9.2(ton N )10.8(W)94.9(.)-1.3( 1995.)]TJ /F4 1 Tf 8.8 0 TD -0.0633 Tw [( Beetle banks )-9.5(\227 helping nature to control)]TJ -7.7921 -1.1087 TD -0.0037 Tc (pests. )Tj /F20 1 Tf 2.5197 0 TD -0.0054 Tc 0.0556 Tw (Pesticide Outlook)Tj /F4 1 Tf 7.1055 0 TD 0 Tc 0 Tw ( )Tj /F19 1 Tf 0.3055 0 TD (6)Tj /F4 1 Tf 0.4945 0 TD -0.006 Tc (:13-17.)Tj /F19 1 Tf -11.4331 -1.1118 TD -0.0049 Tc -0.0134 Tw [(S)48.5(tewar)-4.9(t A)54.9( )-52.7(J A)-5(,)-5( )-39.9(W)23.6(right A)54.9( )-46.6(F)83.3(.)-1.1( )-51.8(1995.)]TJ /F4 1 Tf 13.9055 0 TD -0.0051 Tc 0.0375 Tw [( )48.3(A)54.7( new inexpensive suction)]TJ -12.8976 -1.1118 TD -0.0054 Tc 0.0991 Tw (apparatus for sampling arthropods in grassland. )Tj /F20 1 Tf 19.5937 0 TD -0.0065 Tc 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